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692 :: A: Annals of the Missouri Botanical Garden Figure 3. Diefenbachia beachiana. A. A series of leaves showing adaxial surface (Croat 44283). B. Close-up of leaf cluster showing variegated leaves and a cluster of inflorescences (Sims 8). C. Adaxial surfaces of leaves and open inflorescences. D. Close-up of inflorescence showing emergent portion of the spadix and adaxial surface of petiole. C, D. (Croat 68444).
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia Discussion. The species is characterized by its Cultivated specinzens. Costa Rica. Henny 5 (M0). Hemoderately thin, quilted blades with many broadly redia: Finca La Selva, O.T.S. Field Station, oliginally collectecl by Jim Beach, cultivated at Missouri Botanical Garspreading primary lateral veins and crisped-undu- den (MBG), vouchered as Croat 59152 (M0). late margins with the lower midrib and primary lateral veins puberulent. DieJJ:7enbachia beachiana is 3. similar to D. galdarnesiae7 a species from central Panama that also has somewhat puberulent midribs on the lower blade sudaces. That species differs in having the pubescence much shorter, merely granular-puberulent, and has fewer than 22 veins per side (vs. 23 to 36 for D. beachiana) that arise from the midrib at a 30°-40° angle (vs. 70°-110°). In addition, Panamanian populations of D. beachiana differ in having their blades usually white- to Dieffenbachia burgeri Croat & Grayum, sp. nov. TYPE: Costa Rica. Puntarenas: Palmar Norte-Panamanian border7 3 km N of turn-off to Rincon, 8°48'39"N, 83°16'18"W, 110 m, 10 Sep. 1996, E B. Croat & D. Hannon 79211 (holotype, MO-5170493!; isotypes, AAU!, B!, CAS!, CR!, COL!, DUKE!, EAP!, F!, GH!, HUA!, INB!, K!, M!, MEXU!, NY!, PMA!, RSA!, US!, VEN!, UB!, WU!). Figures 4, 27B. cream-mottled, whereas those of D. galdarnesiae Planta terrestris, 0.5-1 m; internodia 0.5-2.0 cm longa, are always solid green in color. Dieffenbachia be- 1.2-2.5(3.5) cm diam.; petiolus 13-48 cm longus, vaginatus 1/3_2/3 longitudinis; vagina 8.5-20 cm longa, inequiachiana is also somewhat similar to D. grayurni- lateralis acute ad apicem apex; pars libera 5.3-29.5 cm ana7 but that species differs in having the veins longa; lamina oblongo-lanceolata vel anguste elliptica, merely minutely granular in slender rows on drying, 25-40 cm longa, 5-16 cm lata, nervis primariis lateraliand in having somewhat more ovate and wider bus (13)18-28 utroque, inflorescentia l -7 in quoque axblades. illa; peduneulus 7.5-12 cm longus; spatha 1].5-15(21) cm longa; spadix 9-15.6 cm Songus; piKstilla 21-37. A single collection (Cro(t 6695X) from Verclguas Province in Panama (lifferx in having the primary Herlg, (). S- 1 m tall, sa, milky, sharply foetid; latercll veins curving more prominently towar(l the interno(les glossy to semiglossy, (lrying fre(luently apex than rllatericll frolll eIsewhele ill westeln l'cln- lllatte an(l millutely warty, ()..S-2 ( m long, 1.2anla and (3oStcl Ri( a. In cld(lilion, the pul)es;( en( e 2..5(.-3..5) ( m (lianl., (lark green to }v1a( kis;h green, is nlerely granulclr-l)ul)el ulerlt l ather than pubet u- t. H(lltl/tl(l 966()7 (-olle( te(l clt l,(l Selvcl in Heledica Province in Costcl ltica, ploba})ly repreF;ents a hybrid between D. beachiana an(l 1). (on(illn(l. The specilllen has tfile sliglltly s( abrous petioles an(l the impressed Inajol veins of D. b()el(hieln(l, lxut the blades are mole or less elliptic and dry the colol of D. concinn(l. A*lflltiotl(ll s/)ecilrze/ls S()(M (E()S'I'A 141(CA. Heresflia: llfo leje-llfo Sardinalito, Atl. slo^)e of Vol( tirl 13atva, (,r(l- rllm bX99 (M()); Sat1 Jose-lvlo. Viejo, vi(. of Chilarrlarlle, l 1.6 mi. N of (2aril)larle o, Cro(lt be'.X.58 (}3, Cll, M()); Iq'i1( a La Selva, rlear Rfo l'uetlo Viejo, BurW,rer & Stolze 5754 (CK, 1'^); Zona Plote(tora, N slolles of Vol(an l1arl)a, Rfo Peje-Rfo Gua(imo, Grelyum & KSchltz 3220 (DUK1,). Li- mon: Rio Pa( uare-Quebra(la l)iablo, ea. 2.5 km 1E ol Si- quirres, Grayum 7698 (M0); la. rl'oltugueton Est. Agua Fria, R. Robles 1234 (CR, M0); I, Colombiana Farm, United FIU jt CO, Standley 36840 (US). Puntarenas: Gol- fito, Par. Nac. Esquirlas, Quebrada Gamba, Sanchez 539 (M0). San Jose: Vazquez de Coronado, Braulio Carrillo NP, along hwy. San Jose to Siquilres, trail to Rio Sucio, Croat 7X77l (INB, M0, WU). PANAMA. Bocas del Toro: Chiriqui Lagoon, Water Valley, von Wedel 1438 (M0); lO km SW of Chiriqui Grande, Thompson 4937 (CM); Fortuna Dam-Chiriqui Grande, 7.3 mi. N of bridge over Fortuna Dam, 3.2 mi. N of Continental Divide, Croat & Grayum 60255 (M0); Chiriqui Grande-Fortuna, 7.7 mi. W of Chiriqui Grande, 1.5 mi. W of Punta Pena, Croat & Grayum 60094 (K, M0, SC75); near hwy. to Chiriqui Grande, McPherson 11816 (M0). Veraguas: vic. of Santa Fe, 8 mi. N of Escuela Circlo Alto de Piedia, Croat 66953 (M0, PMA, SCZ, US). 693 Iryirlg (Iark })rown to })Iae k Or yellow-l)rown Or yellow-greerl. I,lq,AVF,% s( attere(l along stenl with areas exose(l ex( e)t neal a)eX; /seliol().s 1.-3-48 ( nl long (averagi ng 26) ( m long), (Iark green, s;u rf:d( e some- W}at rll()ttle(lX weakly r)ale-streake(l neal lease, glossy to s;enliglossy, sheathirlg 1/3 tO 2/3 the length of the yetiole (().S their average }ength); sheath 8.5- 2() ( nl long (averaging 12.6 ( m), sometinles soli(l (reamy white, with margins often drying thin an(l light l)rown, the tiy) inequilaterally ae ute; un- sheathe(l portion 5.3-29.5 c m long (averaging 12.9 cm), variable in cross section, from subterete to C- shape(l or U-shape(l, sharply sulcate to somewhat flattene(l near apex a(laxially, the margins obtuse to acute; I)lade.s oblong-lane volate to narrowly elliptic, 25-40 cm long, 5-16 cm wide (averaging 30.5 x 10.5 cm) 1.9-3.7 times longer than wide (averaging 2.8 times longer than wide), 0 83-1.6 times longer than petiole (averaging 12 1 times longer), slightly inequilateral, one side 0.4-1.4 cm wider than the other side, thinly coriaceous to subcoria- ceous, slightly to moderately bicolorous, gradually long acuminate at apex, narrowly acute to rounded and equilateral or slightly inequilateral at base (one sometimes up to 6 mm higher on the midrib than the other); upper surface solid dark green, matte to weakly glossy (sometimes variegated with pale mot- tling), drying gray-green to dark olive-green; lower surface slightly paler, semiglossy to weakly glossy
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762 , C72 89° 86° 83° 80° 77°
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