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712 Annals of the Missouri Botanical Garden Figure 11. Dieffenbachia grayumiana. A. Habit (Croat 60149). B-D. (Croat 74953). B. Leaf blade adaxial surface. C. Blade abaxial surface. D. Apex of stem showing heavily sheathed petioles with sharply sulcate free portion, and speckled petiole bases.
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia from January to June, rarely as eclrly as November. rante-13ocas del Toro, near Milla 5, Croat & Porter 16499 Fruiting collections have been seen only from May (MO); Chiriqui Grande-Fortuna, 3 mi. W of Chiriquf Grande, Croat & Grayum 60149 (K, MO, PMA, US)7 Gualto September. aca-Chiriqui Grande, 8.1 mi. S of Punta Pena, Croat 74953 Discussion. The species is characterized by its (MO, PMA). Chiriqui: Gualaca-Chiriqui Grande, 1.4 mi. S narrowly ovate, typically subcordate, mottled blades; weakly sheathed, decurrent petioles; and of Punta Pena, Croat 74945 (MO), M. Akers 78A (MO). variegated stems and petioles. Also characteristic 12. Dieffenbachia halnlnelii Croat & Grayum, are blades that are frequently glossy on the upper Novon 9: 496. 1999. TYPE: Costa Rica Hesurface and matte or nearly so on the lower surface. redia: Finca La Selva, O.T.S. Field Station on The major veins are sometimes minutely farinose- the Rio Sarapiqui, 50-80 m, M. H. Grayum granular on the lower sudace. In this regard it is 7670 (holotype, M0-3491533!; isotypes, B!, similar to D. beachiana: a species with puberulent CR!, K!). Figures 12, 27B. major veins on the lower blade surface. Aside from pubescence type, D. beachiana also differs in hav- Small herb, 25-40(70) cm tall; sap not fouling much narrower blades (1.8-5.3 times longer smelling; stem becoming decumbent and subrhithan wide) with more numerous pairs of primary zomatous at base; internodes 1-1.5(2.7) cm long, lateral veins (23 to 36 pairs). on lower portions, 3.5-7.5 cm long toward the apex, Dieffenbachia grayumiana is superficially simi- 0.5-2 cm diam., glossy, dark blackish green, drying lar to D. seguine from the West Indies in the shape yellowish brown to gray-green, weakly striate. and coloration of its dried blades but lacks the LEAVES erect-arching; petioles 7-25(35) cm long sharply sulcate petioles, the bicclrpellclte ovaries, (averaging 15 em long), erect, green or mottled with and the protruding stubby spadix with cl reflexed dull yellow-gl efen, (lrying greenish to sometimes spclthe b]cl(le seen in the Lltter. yCll()WiSh br()Wn, XUltd('8 (11 ying mdttC, XhUdthing A ( olle( tion from BO( clS (Je1 r1^oro Provine e in f()r (().
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