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754 Annals of the Missouri Botanical Garden km upstream from and NW of Puente Boquer6n, 8.6 mi. throughout (this white coloration continuing onto SW of Catacamas, 6 mi. SW of Sta. Maria del Real, Croat the midrib), sheathed virtually throughout; sheath & D. Hannon 64109 (INB, MEXU, MO, TEFH, US); along Rio Olancho, on rd. Gualaco-San Esteban at Rio San erect to involute (rolled inward throughout in age), Martin, 19.3 mi. E from Gualaco, 8.7 mi. SW of San Es- free-ending and unequally rounded at apex, proteban, Croat & D. Hannon 64317 (B, MO, US); Rio Olan- longed to 2 cm beyond the base of blade; uncho, San Esteban-Bonito Oriental, 19 mi. NE of San Es- sheathed part obsolete or rarely to 1 cm long (when teban, Croat & D. Hannon 64476 (MO); Refugio de Vida Silvestre "La Muralla," Nelson & Andino 16247 (TEFH); evident obtusely flattened); blades ovate to ovate- Mpio. Jano, 16 mi. NE of La Union along the rd. to Olan- elliptic or oblanceolate-elliptic, (15-)25-48(-63) hito, Davidse et. al. 35484 (MO). Yoro: near Puente cm long, (8-)15-32 cm wide (averaging 34 x 16 Grande, on a tributary of the Rio Agua (Rio Puente cm), broadest near the middle, sometimes below, Glande), Blackmore & Chorley4073 (MO). NICARAGUA. frequently above the middle, 1.4-2.5(-3.5) times Matagalpa: El Tollsa Rd., Neill 1571 (MO); ridge along td. betw. fJa Dalsta arld La Luna, E of Esquipulas, Stevens longer than wide (averaging 2.1 times longer than 11786 (CAS, M() TEX); Macizos de Penas Blancas, SE wide), 1.9-3 times longer than petioles, spreading side, drainage ol (ssuebrada E1 (ssuebradon, Hda. San Mar- to erect-spreading, inequilateral, one side 1-3 cm tin, horder with l)epartmento de Jinotega, Stevens et al. wider than the other, gra(lually to abruptly acumi- 2()998 (MO); Matagalpa-Siuna, 1.5 km al NE de Los Angeles, Moreno 17/42 (MO); E1 Trebol, 7 km S of Penas nate, sometimes acute at apex, inequilaterally cor- T3lancas, rd. to fi l 'luma, Moreno & Robleto 20526 (MO). dulate at base, one side sometimes broadly rounded Zelaya: Finca Waylawas, Pipoly 4479 (MO, US); Siuna- to obtuse, the other side cordulate, sometimes in- Matagalpa, ca. 31.4 km beyond Rio Uli (near Wani), ca. equilaterally acute, subcoriaceous, often conspicu- 8.9 km beyond llosa Grande La Balsama, Stevens 7457 ously quilted, moderately l)icolorous; upper surface (MEXU! MO); Cerro l ivico, 7 km NE of Siuna, Neill 3629 (MO); Rio Sucio, Ed of Bonanza, Stevens 12311 (MO), Ste- usually solid dark to medium green, sometimes vens 8044 (MO); Res. Bosawas, Bonanza, Cerro Cola Blan- conspicuously to sparsely variegated with pale ea, vic. Cacerio (le Vitinia, Rueda & Coronado 6557 (MO). green or pale yellow thloughout much of the surface, the mottling large ol small, but somewhat re- 25. Dieffenbachia tonduzii Croat & Grayum, stricted to the area midwcly t)etween the midrib and Novon 9: 497. 1999. TYPE: Panama: Bocas del margin, matte to weak]y glossy, sometimes appear- Toro: Valle del Silencio, along Rio Changuinola, ing weakly velvety, drying glay-green to olive-green ca. 1 km above mouth of Rio Teribe, vic. Teribe or dark brown; lower surfcl( e much paler and matte Indian population, disturbed forest among co- to weakly glossy, silvely-gleen, drying yellowish coa plantations, 9°21 '40"N, 82°31 '40"W, less green to yellow-brown helow; niidrib flat to broadly than 100 m, 25 June 1994, 1: B. Croat & G. rounded and moderately to strongly paler, pale Zhu 76452 (holotype, MO-04611212!; isotypes, green or sometimes creanly white above (sometimes AAU!, B!, BM!, BR!, CAS!, CM!, COL!, CR!, only toward the apex), blutly acute to obtusely an- DUKE!, F!, GH!, HUA!, INB!, ITIC!, JAUM!, gular and paler, sometillles white or creamy white K!, L!, LE!, M!, MEXU!, NY!, P!, PMA!, below, (0.6-)1-1.7 cm wi(Se; priniary lateral veins QCA!, QCNE!, R!, RSA!, S!, SCZ!, SEL!, (14 to)l8 to 25(to 30) per side, arising at an acute TEX!, UB!, US!, VEN!, WU!). Figures 25, angle and spreading at 45°-90°, sometimes reflexed 29B. toward the base, promillently to weakly and obtusely sunken above, convex to weakly raised and Terrestrial herb, 0.5-1.5 m tall, usually to less darker than surface or concolorous below, some of than 1 m tall; internodes 1.5-4.5(-6) cm long, 1.5- the lowermost with a weak fold near the base (Croat 3(-4.5) cm diam., usually solid dark to medium & GrayunI 60112), somelillles convex-pleated begreen, sometimes faintly marbled with gray-green low; interprimary veins almost as conspicuous as or yellowish gray throughout (on plants that also the primaries; minor veins moderately to distinctly have streaked petioles), initially weakly glossy, be- visible, darker than surface below. INFLOREScoming semiglossy to glossy, often with a subvelvety CENCES 1 to 2(to 4) per axil, often with two orisheen; petiole scars manila to white, curved down- ented in opposite directions; peduncle (3-)6-17 cm ward on the opposite side of the stem and ending long (averaging 10.3 cm long), 7-8 mm diam., unevenly; petioles 10-24 cm long (averaging 17.6 weakly glossy, dark to medium green, sometimes cm long), held + erect, medium green (except with pale yellow-green streaks; spathe (12-)15-28 sometimes white to pale green at base), almost cm long (averaging 20 cm long), 2-4 times as long matte to weakly glossy, weakly striate (especially as peduncle, acuminate at apex, convolute to about near the base), narrowly rounded to obtusely an- the middle in lower part, matte to weakly glossy gular on abaxial surface and often white medially, outside, glossy within, solid medium green on both sometimes streaked in a variegated pattern surfaces, gradually and weakly constricted some-
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia Figure 25. Diegenbachia tonduzii. A. Close-up of stem, showing both creeping and erect portions. B. Habit with inflorescences and variegated blades. C. Crown of plant with open inflorescence. D. Close-up of stem showing petioles streaked with creamy white. E. Close-up of stem showing solid green petioles. F. Close-up of spathe with the blade portion open (Croat & Zhu 76452¢ type plant). G. Inflorescence open to show upper male flowers and lower portion of spadix with pistils and staminodia (Croat 66533). A, B, C. (Croat & Zhu 76452¢ type plant); D, E. (Croat 76450). 755
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