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704 r Annals of the Missouri Botanical Garden Figure 8. Dieffenbachia davidsei. A. Potted plant showing habit with several inflorescences. B. Blade adaxial surface with quilted primary lateral veins. C. Close-up of stem showing mottled petiole and stems. D. Crown of plant with cluster of inflorescences, and petiole showing free-ending sheath apex. E. Abaxial surface of leaf blade and immature inflorescence. F. Close-up of open inflorescence.
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia clbove, convex and paler than surface below, fre- The species frequently occurs in mature forest quently pale-mottled; primary lateral veins (6)9 to along streams. 15 per side, flat to sunken7 weakly quilted above, Phenology. lliefienb(zchia daridsei flowers in raised and darker than surface below, drying usu- the rainy season sometime after May (one plant ally darker than surface below, sometimes paler seen with both flower buds and seemingly mature than surface, usually departing midrib at 65°-90° infRorescences in lclte October). Mature fruits have angle throughout most of its length (rarely with the been seen during the dry season, in the late rainy veins in the distal half of the blade arising at 30°- season? and early dry season from November 50°), prominently curved toward the apex in the through April. Immature fruits have been seen in outer 1/3, those near the base to as much as 120° late April indicating that flowering may begin even clngle, sometimes forming a weak sigmoid curve, earlier than May. the primary lateral veins in the upper 2/3 to 1/2 some- Discussion. The species is distinguished by its times loop-connected for some distance, forming a slender, dark, semiglossy, mottled stemsS mottled weak collective vein 2-5 mm from margin, but this petioles with a prolonged, usually acute sheath usually not extending all the way to the apex; in- apex, and more or less oblong blades with rather terprimary veins almost as prominent as primary prominent primary lateral veins that spread at nearlateral veins, flat to sunken above, weakly raised ly 90°. and darker than surface below. INFLORESCEN- Dieffenbachia davidsei is probably most closely CES usually to 3 per axil, sometimes with more related to D. obscurinervia. That species differs in than one axil producing inflorescen(es; /eduncles usucllly being cl larger plant (to 1.5 m) with plom- 1.5-4 em long, 4-5 mm (tiam., ovoi(l to D-shape(l inently scurfy brown internodes, petiole sheclths in c.ross se( tion; spathe 8-16 cm long, 3.1.-4.8 which clre tyl)i(cllly only rounde(l clt the clpeX (not times longer than pedunc le, e oria( eous! semig]ossy free-en(ling), cln(l [)lclflex which hclve weclkly (leve,lon hoth sulfates, me(lium lo light gleen outsi(le, ol)e(l I)rimclly lcltelcll veins thcltcllisecltcln angle, slightly 1)a1e1 within; tuT)e olulol, lo someewhcll el- tlstlcllly cll)otlt 4.5° lo lle llli(lril). lil.)soi(l, G-7 cm long, ol)lo]g-lfll( volclle wherl All.[o|g[ l[]e [)IbOa(lly sI)Xea(linn l)l'illlal'y lcllPldl ol)ele(l, a( llle at apex (a(+lillell tIl)i( lllcilP ['01' 2 rilill); VEillE t11P 011P 0t lllP IllOSl (liSlill('liVP ('llcil('lPliSli(bE .Nvelelit (7.5)1()-12.5 e m loly,* uI) lo 1.7 em shollet 0t' 1). el(llpi(l..sei (willl llle VEill.S [lELlcllly S[)l'vtl(lillS lhan the s^)clthe; pistillate ^)oltioll (4.()).5..5-() ( m fr0nl lilP nli(lril) cit a l)lOcl(l AnglC PVEI1 to nEcil tilC long, (())tS-I() rnm wi(le (the clXiS ('a. 5 t11l11 (licllll.)t A})UX 01 tilP 1)ltl(lU), cl fEw ('OllU('tiOllE [tOtil thP ,1 alrrlosl ( olltig^.lous with the stalrlill.lle l)oltioll (a I ltino-Carli ltocl(l in l'cinatila (est)e( icilly lhe I'MA wi(le stelile illtermeeticlte se(liorl lclekirg); the l)is- XllUPt 01 I(lre(le.s 94()) hcive t)rinicily lciteral veinx tillclle [IOWEI S more wi(tely St)ct( C(l llUcil tilC cl[)eXe lhat xl)tel(J cil atlgles of u^) lo 4tS°. 'lnhe lvelreele.s 94() 4-6 rrlnl cll)cllt; l)istils 2()-.-J.S, ovaly glol)ose, ^)ale olle( tiol] alKso 11as t1101'e (001StVi( uouSly [)unt lEllC gleen; stiglna l)right yellow with 10llr l)l ushy l)al)il- ^)etiolex tharl (Jovs lle M() sheet of ll]C SalnC nUtIIlae, 2-2.4 Illnl diam.; stamino(lia whiteX usuall.y bel. It iXs l)ossil)le that l'(lre(le.s 94() is a hyl)li(l bet)roa(lly unite(l at base an(l tolming an often nearly tween l). delviel ei al(l I). ol).scllrinel-vi(l. IXoth sI)ec onlplete bowl around the l)istil, somewhat flattene(l ('ies OC('UI in the Nusigandf area along the FJ1 throughout, barely thickene(l at the a,oex, 3.4-5 mm 1,lano-Calti Road. Dieffent)(lchia ol)sezlrinervia also long; stam inate portion oblong 4-6 c m long, has c onspic uously mottled petioles, but its veins creamy white, bluntly pointed at apex; synandria arise at a much more ae ute angle. The stems of D. irregularly 4- to 6-lobed, drying 1.5-2.3 mm diam., obsezlrinervia differ markedly in being matte and deeply depressed with the margins turned up. IN- scurfy-brown rathel than semiglossy, dark black- FRUCTESCENCES with spadix 5.5-7 cm lc)ng, green with yellow-green mottling as in D. davidsei. berries orange, 8-ll mm diam., subglobose, 2- to Another unusual vollection, possibly also a hybrid, 3-seeded. is Croat & Zhu 76666 from La Mesa in Cocle Province near E1 Valle de Anton. It has the glossy in- Distribution and habitat. DieJ%enbachia david- ternodes of D. davidsei but the veins of D. obscusei ranges from northeastern Costa Rica to Panama rinervia. It is perhaps just an unusual collection of (Darien Province) and Colombia (Choco) in Pre- D. obscurinervia. montane wet fnorest (P-wf), Tropical wet forest (T-wf), Etymology. The species was first collected in and Premontane rain fnorest (P-rf) life zones (Hold- Panama by Scott Mori in 1974 along the E1 Llanoridge, 1967). In Costa Rica it is known only from Carti road (Mori et al. 4184), and later in Costa the Sixaola region at 20-40 m elevation. In Panama Rica by Gerrit Davidse and C. H. Hamilton (Davit ranges from 100 to 900 m in Panama Province. idse & Hamilton 23617) in 1983. It is named in 705
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760 Ez D. fosten (0, -8 ' ; < * ( \
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762 , C72 89° 86° 83° 80° 77°
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