l - CNCFlora

700
Annals of the
Missouri Botanical Garden
brown, minutely granular to somewhat scabridu- free section to 1.1 cm long in D. copensis). Dieffenlous;
blades narrowly oblanceolate, 25.5-33.3 cm bachia galdamesiae differs in having blades prolong,
5.2 cm wide, 6.4 times longer than wide, nar- portionately broader, 2.7-4.2 times longer than
rowly long-acuminate at apex, narrowly attenuate at wide, versus 6.4 times longer than wide in D. cobase,
dark green and matte above, moderately paler
and semiglossy below, drying dark gray-green and
penszs.
minutely granular above, light yellowish gray be- 6. Dieffenbachia crebripistillata Croat, sp. nov.
low; midrib concolorous and narrowly raised above, TYPE: Panama. Cocle: Alto Calvario, above E1
eonvex and granular-puberulent to hispidulous and Cope, ca. 6 km N of E1 Cope, Atlantic slope
slightly darker below; primary lateral veins 9 to 12 along trail which leads down to Las Ricas, Lipairs,
arising at an acute angle then spreading at mon 8 San Juan, 8°39'S, 80°3C'GT, 710-800
45°-50° angle, plominently curved upward along m, 22 June 1988, YS B. Croat 68746 (holotype,
the margins ancl extending to above the origin of MO-3610884!; isotypes, B!, (2AS!, COL!,
the next higher veins, paler and weakly sunken DUIkE!, F!, GH!, HUA!, INB!, K!, M!, MEXU!,
above, convex an(l slightly darker below, drying SY!, PMA!, US!). Figures 7, 27A.
narrowly raise(l and granular; minor lateral veins
weakly visible; c ross-veins equally visible on lower Planta 30-100(120) cm alta; interno(libl 2.5-4 cm longa,
1.5-3(5) cm diam.; petiolus 7.5-24 ( lls longus; lamina
surface; lower surt:ace moderately granular. INFL0ovata
vel raro elliptica vel oblongo-elliptica, (22)27-
RESCENCE sol i lary, longer than the petioles; pe- 46(55) cm longa, 10-25 (m lata, nelvis primariis lateralduncles
5 cm long (Irying 2.5 mm diam., pale yel- ilous (10)15-17 utroque; inllorescentis 1-2 per axillam;
low-brown; sp(lth() 1 5.T cm long, 1.2 cm diam., pale peedunculus (8.5)10-14 cm longus; sl)alls{l 13-28 cm longreen,
clrying me(liurrl yellow-brown, narrowly long- ga, 2.0-3.2 cm lata; spadix 13-26.5 ( lls losgus; parte pistillata
6-10(12) cm longa, 8-12 mm lll.
acuminate at tlle tal)ex (acumen 1.7 cm long); spadix
12.3 cm long; fle e portion 8.5 cm long; pistillate Short, thick-stemme(l herb, 3()-1()()(120) cm tall;
portion S cm lollg. 4-S mm diam.; staminate portion sap foul-smelling, eventually tulnilltg white; inter-
5.5 cm long, z1 lllrll cliam., drying yellow-brown; nodes clark green, glossy, 2.5-4 ( lll long, 1.5-3(5)
mostly nake(1; illte.rmediate sterile portion 1.1 cm (m diam.; petio1Kes 7..S-24 cm long (averaging 16
long, 2.S mm (lialll. on drying, with only a few sta- e m long), somewhat s)ongy, soli(l (ltark green or
minodia scattere(t throughout its length; pistils 38, -with a white band on the abaxial surface, conspic-
2 to 3 situated across the width of the spadix; ovary uously sheathed ranging from 0.7 to fully through-
0.5-6.0 mm (liarll.; stigmas about as broad as the out the length (but rarely ending mole than 1 cm
ovary; stamino(lia 2 to 4 per pistil, 1-1.4 mm long, lrelow the blade and often extending beyond the
slightly thickene(t toward the apex, not at all broad- l)ase of the blade); sheath to 1.5 cm high, 7.5-19.5
ened toward the l)ase; the sterile male flowers in 2 em long, inequilaterally rounde(l to auriculate at
rows, somewhal separated from the fertile flowers; apex; unsheathed portion usually not apparent but
synandria 2.2-2.tA mm long, 1.6-2.0 mm wide, ir- when present 1-7 cm long (averagmg 1.6 cm long),
regularly rounded to rhombic at apex. INFRUC- t)roadly sulcate, the margins shalp; blades ovate to
TESCENCE not seen.
larely elliptic or oblong-elliptic, (22)27-46(55) cm
long, 10-25 cm wide (averaging 37 x 18 cm), 1.5-
Distribution (I nd habitat. Dieffenbachia copen- 3.3 times longer than wide (avelaging 1.9 times
sis is known only from the type specimen in the longer), 1.9-3.7 times longer than petioles (aver-
Cocle Province of Panama at 700 to 900 m in the aging 2.3 times longer than petiole)* somewhat in-
Lower montane rain forest (LM-rf) life zone (Hold- equilateral, one side up to 0.5-2.7 crll wider, modridge,
1967).
erately coriaceous, abruptly ac u minate and
Discussion. The species is characterized by its downturned at apex, obtuse to rounde(l at base, the
narrowly oblanceolate, narrowly long-acuminate basal portion of the blades held solllewhat erect;
blades with densely granular primary lateral veins upper surface glossy to moderately glossy, usually
and a gl anular-hispidulous midrib on the lower sur- solid dark green sometimes mottled with pale
face. The species is somewhat similar to both D. green, slightly bicolorous; lower surface glossy, palgaldamesiae
and D. fortunensis, differing from both er; midrib flat-raised to flat-rounded and paler esin
having proportionately narrower blades that are pecially near base, 12-20 mm wide, becoming obbroader
well above the middle. In contrast, the tusely convex toward apex and diminishing before
blades of D. fortunensis are broadest well below the reaching the apex above, broadly flat-rounded to
middle and have the pistillate and staminate por- narrowly rounded and pale green to white below
tions of the spadix contiguous or nearly so (vs. a (especially near the base); primary lateral veins