l - CNCFlora

706 Annals of the
Missouri Botanical Garden
honor of Gerrit Davidse of the Missouri Botanical base, minutely granular-roughened throughout with
Garden, co-editor of the Flora Mesoamericana se- white raphide cells; unsheathed portion 3-15.5 cm
ries.
long, sharply C-shaped, flat-sulcate, with margins
Paratypes. COSTA RICA. Limon: Quebrada Mata de
Limon, Finca Anai (Sixaola region), Grayum fst al. 4440
(CR, MO). PANAMA. Bocas del Toro: Fortuna Lake
Area, Fortuna-Chiriquf Grande, 1.6 mi. N of Cont. Divide,
prominently raised; blades narrowly oblong-lanceolate,
rarely sometimes narrowly ovate, or narrowly
oblanceolate, 15-33 cm long, (3.5)4.5-9.5(13) cm
wide (averaging 21 x 6.8 cm), 2-6.3 times longer
Croat & Zhu 76533 (MO). Cocle: Cano Blanco del Norte-
Cano Sucio, Davidse & Hamilton 23617 (MO, PMA); E1
Cope, Cont. Divide, Hammel 13685 (MO); E1 (:ope, ca. 1/2
mi. N of Cont. Divide at Alto Calvario, Croat 75100 (MO);
Atl. slope, Hammel 5607 (MO). Darien: Par. Nac. del
Darien, of Rfo Pucuro, vic. Tacarcuna, ca. 18 km E of
Pucuro, Hammel et al. 16397 (MO). Panama: E1 Llano-
Cartf Rd., 9.8 km from Interam. Hwy., Mori et al. 4184
(MO). San Blas: trail to Cerro Camucanala from Rfo Titamibe,
de Nevers et al. 4709 (MO, PAN); (:omarca de
Kunayala, Nusigandl, E1 Llano-Cartf Rd., 10.1 mi. N of
than wide, 0.38-1.48 times longer than petiole (averaging
1.1 times longer), + equilateral or weakly
inequilateral, one side 5-15 mm wider than the
other side, thinly coriaceous to sulgcoriaceous, drying
papyraceous, usually quilted, weakly lgicolorous,
+ equilaterally or sometimes inequilaterally
acuminate or sometimes acute at apex (acumen
sometimes lgriefly apiculate), equilateral to slightly
inequilateral an(l attenuate to acute, rarely rounded
Interam. Hwy., Paseo Mariska, Croat & Zhu 77017 (INB, at lgase; margins usually conspicuously undulate,
MO); E1 Llano-Cartf Rd., de Nevers & H. Herrera 3975 drying crisped; upper surface glossy to semiglossy
(MO); km 18, Galdames et al. 1330 (STRI); Nusigandi, E1
or weakly glossy, medium to dark green, sometimes
Llano-Cartf Rd., betw. Nusigandf and 1 mi. N of Nusigandi,
Croat & Zhu 76597 (HUA, MO); Sede de Campo mottled white or cream; lower surface matte to
de PEMASKY, ca. 2() km on E1 Llano-Cartf Rd., Paredes weakly glossy, mo(lerately paler; m.idril) flat or flatet
al. 940 (MO, STRI); Isla Nargana, tributary of Rfo Dia- raised and concolorous above, thicker than broad
blo, Galdames et al. 1570 (PMA, STRI). Veraguas: Cerro
Tute, above Alto de l'iedra, McPherson 10725
to convex below drying lgrownish matte and darker
(MO). CO-
LOMBIA. Choco: Nuquf, Corr. Arusf, Est. Biol. E1 Amar- than surface to slightly paler than surface, granulargal,
along trail to Arusl, Croat & Mor(l 83685 (MO). puberulent to conspicuously granular below; primary
lateral veins 1() to 17 per side, departing mid-
8. Dieffenbachia fortunensis Croat, sp. nov. rib at an actlle angle, then moderately curved
TYPE: Panama. Chiriquf: along road between toward the margin at 55°-90° (mostly 55° in the
Fortuna Lake and Chiriquf Grande, 4.5-5 km middle of the }lade but at more acute angles toward
N of dam of Fortuna Lake, 8°43'N, 82°17'W, the apex and more olgtuse angles toward the lgase),
1100-1135 m, 8 Mar. 1985, E B. Croat & M. turning upwar(l along the margins and forming a
H. Grayum 60002 (holotype, M0-349012!; iso- series of fine collective veins along the margins,
type, PMA!). Figures 9, 28A.
some of which extend to the apex, matte, prominently
sunken to narrowly or weakly sunken and
Herba 4s100 cm alta; internodia 1-5 cm longa, 0 8- concolorous at)ove, thicker than broad or convex
2.0 cm diam., petiolus 12-28 cm longus, vaginatus 0.37-
0.83 longitudinis; vagina 6.5-14.5 cm longa, decurrens ad
apicem; lamina oblongo-lanceolata, raro interdum anguste
ovata vel anguste oblanceolata, 15-33 cm longa, 4.5-9.5
and darker than surface, granular-puberulent to conspicuously
granular below, drying darker than surface,
interprimary veins usually about as conspicucm
lata, nervis primariis lateralibus 10-17 lltroque; inflo- ous as primary lateral veins, minor veins moderately
rescentia solitaria;
12-18 cm longa;
pedunculus
spadix 9-11.5
4.7-7 em
cm longus.
longus; spatha
obscure to distinct; "cross-veins" loose-connected,
weakly oblique, drying minutely dark-punctate. IN-
Slender herb, 40-100 cm tall, sap lacking dis- FLORESCENCFJS 1 per axil, peduncle 4.5-7 cm
tinctly foul aroma, stem erect except at base, con- long, spathe 12-18 cm long, 2-3.7 longer than spataining
inconspicuous, mThite raphide cells, inter- dix, 1-1.5 cm diam. on tule when furled, gradually
nodes 1-5 cm long, 0.8-2 cm diam., unmarked, contracted midway, gradually long-tapered to apex,
semiglossy, dark green to medium green, sometimes uniformly pale green on both surfaces, spadix 9-
minutely mTrinkled and glistening, drying black- 11.5 cm long, drying 3-3.2 mm diam., free portion
ened, moderately smooth (lzut minutely granular on 1.5-2 cm long, drying 2 mm diam., pistillate pormagnification).
LEAVES erect-spreading, + clus- tion 4.24 cm long, drying 7 mm diam. throughout,
tered at/near stem apex, petioles 12-28 cm long fertile staminate portion 54.5 cm long, drying 4
(averaging 19.6 em long) sheathing for 0.37-0.83 mm diam. throughout, mostly naked intermediate
their length (averaging 0.58 their length), sheath portion of spadix 1.5-2.0 em long, drying 2 mm
6.5-14.5 cm long (averagin; 10.1 cm), with mar- diam., with a few sterile pistillate flowers in the
gins striate, decurrent distally, dark green, drying lower half, pistils 35 to 38, well-spaced, rarely
blackened to dark bromTn, surface matte, striate at more than 2 dispersed across spadix width, de-