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746 Annals of the Missouri Botanical Garden longer than petioles, shortly acute and markedly a broad sterile segment rather than having them inequilateral above, acute to obtuse at base, pri- contiguous or nearly so. mary lateral veins 6 to 7 per side, concentrating mostly in lower half of the blade, extending parallel 23. Dieffenbachia seguine (Jacq.) Schott, Wieto the midrib, then spreading at 25°-30° angle, ner /. Kunst 1829(3): 803. 1829. Arum seguinearly straight to weakly curved to the margin and ne Jacq., Enum. Syst. P1. 30. 1760. Arum sesweeping up along the margin, drying moderately guinum, orth. var. Caladium seguinum (Jacq.) obseure above, convex and darker than surface be- Vent., Mag. Encycl. 30. 1801. Diegenbachia low; minor veins parallel with obscure and irregular plumieri Schott, Oesterr. Bot. Wochenbl. 69. ross-veining on drying; surface drying matte, mod- 1852. TYPE: Plumier, Descr. P1. Amer. tab. el-ately dark yellow-brown above, light brown and 61 (lectotype, designated here). Figures 23, finely pale-speckled below. INFLORESCENCES 2 30B. pel axil; peduncles 3.5-5 cm long, drying light CaladiunI niaculatunI Lodd., Bot. Cab. tal. 608. 1822. brown, 2 mm diam.; spathe 17 cm long, to 3.5 cm wide when flattened; spadix 14.5 cm long; pistillate portion of spadix to 8.5 cm long, 4 mm wide; staminate portion of spadix 5 cm long; mostly sterile Dieffenbachia niaculata (Lodd.) G. l)on, in Sweet, Hort. Brit. ed. 3, 632. 1839. Dieffenbflchia niaculata (Lodd.) Bunting, Baileya 10: 145. l 963, nom. superfl. TYPE: Lodd., Bot. Cab. tab. 6()8. 1822 (lectotype, designated here). segment 2.2 cm long, drying 2 mm diam., with a few scattered cup-like pistillodes throughout its length; pistils 59, borne in a cup-like disk; staminodia mostly missing. Dic)Jfenbachia literata Schott, Oesterr. Bol. Wochenbl. 68. 1852. Dieffenbachia seguine fo. litllrata (Schott) Engl., F1. Bras. 3(2): 175. 1878. I)i(.ffenbachia seguine subvar. Iiturata (Schott) lErlr,l., Bot. Jahrb. Syst. 26: 568. 1899. Dieffenbachi(l .s(guine var. Iiturata (Schott) Engl., Pflanzenr. 1V, 2.< Dc(Heft 64): Distribution and habitat. Dieffenbachifl pittieri is known only from the type in the Isthmux of Panama in an area of Tropical moist forest (T-mf) life zone (Holdridge, 1967) at less than 200 m elevation. It is most unusual to encounter an endemic species in this life zone, but the species is distinct from other species in Dieffenbachia. There is a possibility that this could be a hybrid between D. kil- 47. 1915. TYPE: based on a cullivtlted collection fide Schott (not seen). Schott Pailllillr 1874 serves as the type [microfiche 64. c T] (I( ( lolype, designated here). I)ietfienbachia lineata K. Koch 8 BoLlel>, Index Sem. (Berlin) 14. 1853. Dieffenbachifl seglline fo. Iineata (K. Koch & Bouche) Engl., F1. Blas. :s(2): 175. 1878. Dieffenbachia seguine subvar. Iill(ata (Schott) Engl., Bot. Jahrb. Syst. 26: 569. 1899. Dicwffenbachia seguine var. Iiturata (Schott) Engl., Pflanzenr. IV, 23 lipii and D. isthmia because hybridization does oc- Dc(Heft 64): 47. 1915. TYPE: New Cranada, based c ur in the wild in Dieffenbachia. Its greatest on a cultivated collection at the Bf rlin Botanical Garden, not sczen. Schott illustratioll 1867 serves as esemblance is with D. killipiiS which also occurs the type [microfiche 64 d T] (le(lol!|)e, designated in this area and dries a similar color. Dieffnenbachia here). pittieri differs from D. killipii in having a dark Di().ffenbachia robusta Schott, Oesterr. I3OI. Wochenbl. 65. brown, smooth-drying stem (light yellow-brown, 1854. Dieffenbachia seguine var. robllsta (K. Koch) Engl., Bot. Jahrb. Syst. 26: 568. 1Xf39. TYPE: Lousually conspicuously ridged stems for D. killipii), cality unknown, cultivatczd colle( tioll of unknown orfully sheathed petioles (usually with the petiole igin at Berlin Botanical Garden. S( hott illustrations sheath ending well below the base of the blade for 1907-1908 serve as the type [mi( I-ofi( he 66 b2, b3] D. killipii), blades with the midrib flat on the upper (lectotype, designated here). Dicvffenbachia consobrina Schott, Syn. Aroid. 131. 1856. surface ("square-raised" for D. killipSi), and a spa- TYPE: Brazil. Rio Negro, Martius s. rl . (holotype, M) . dix with the pistillate and staminate portions widely Dieffenbachia poeppigii Schott, Syn. A Z oi(l. 130. 1856. separated by a long, mostly sterile portion (vs. with TYPE: Peru. Without locality, Poep/)ig s.n. (holotype, the pistillate and staminate portions more or less W destroyed during World War II); S( hott illustration 1903 serves as the type (microfiche 64. b 6) (lectocontiguous in D. killipii). type, designated herc). Dieffenbachia pittieri has been confused with D. Dieffenbachia cognata Schott, Syn. Aroid. 130. 1856. Obscurinervia (Standley, 1944; Croat, 1978), but it TYPE Suriname. Paramaibo, I}Iartius s.n. (holotype, proved to share little in common with that species M). [See also Schott paintings 1845 & 1846.] Di().ffenb(lchi(l gollnterian(l Sf hott, Oesterl. Bot. /. 8: 387. once the type specimen was examined. It differs l858. rl=YPE: Venezuela. Without lo( ality, Collmer from that species in having smooth, not scurfy s.n. (holotype, B apparently lost); Sf hott drawing stems, unspotted, fully sheathed petioles, blades 1857 (microfiche 64 b. T) serves as the type (lectowith distinct primary lateral veins and with a midtype, designated here). Die.JJenbachia neglecta Schott, Bonplandia T: 30. 1859. rib flat, not "square-raised" above, and a spadix TYPFi: Jamaica. Without localityS Distin (holotype, with pistillate and staminate portions separated by K!).
Volume 91 Number 4 2004 Croat Revision of Dieffenbachia Figure 23. Dieffenbachia seguine. A. Habit of plant showing the long creeping caudex and erect portion of stem apex (Croat 60613). B. Potted plant showing habit and leaves with variegated blades and inflorescences at anthesis (Croat 69695). C. Close-up of crown of plant with abaxial surfaces of blades and side view of the spathe (Croat 53921). D. Petiole showing slightly free-ending, inequilateral sheath and acutely sulcate free portion above sheath (Croat 68494). E. Crown of plant showing speckled petioles and inflorescence at anthesis (Croat 77298). F. Spathe at anthesis, cut open to show inner surface of tube. G. Close-up of inflorescence showing apical half with spathe blade somewhat spreading and staminate portion of spadix protruding somewhat forward. H. Portion of spadix showing bicarpellate pistils. F, G, H. (Croat 71828). 747
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