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718 Annals of the Missouri Botanical Garden between D. horichii (Grayum 4756) and D. oerstedii sheath 8.5-21 cm long (averaging 14 cm), with (Grayum 4765). It differs in having petioles that are margins drying thin, light brown and minutely unnarrower and less fully sheathed. dulate, the tip inequilaterally acute to emarginate Etymology. The species is named in honor of and free-ending; unsheathed portion flattened or horticulturist Clarence Horich, who made the first rounded and becoming weakly sulcate toward apex collection of the species. in cross section (never sharply sulcate), blunt to Paratypes. COSTA RICA. Puntarenas: San Jose Province, Playa Dominical-San Isidro del General Baru, 'rinamastes, Burger et al. 10669 (F, MO), Burger & Baker 10137 (CR, F), along Quebrada Bonita, Carara Res., Grayum et al. 5723 (INB, MO), Finca E1 Eden, km 183, R. 2, ca. 400 m E of Santa Marta, L. G6mez 22951 (B, CM, CR, MO); Quebrada Bonita, Carara Res., Grayun 4756 (CR, MO), Croat 79073 (EAP, INB, MO, PMA), hills at SW part of Montanas Jamaica, ca. 2.5 km NE of Bijagual moderately acute, rarely broadly and bluntly sulcate; blades oblong-ovate to narrowly ovate, (12)15- 30(39) cm long, (6)10-26 cm wide (averaging 26 x 16 cm), 0.78-1.77 times longer than petiole (averaging 1.1 times longer), inequilateral, one side 0.5-2.6 cm wider than the other side thinly coriaceous, abruptly to gradually acuminate at apex, acute to cordate at base, the sides often + unequal, de Turrubares, Carara Res., Grayu1m et al. 5467 (MO). usually subcordate with at least one side subcor- San Jose: Par. Nac. Braulio Carrillo, Quebrada Sanguijuela, Chavarrfa & Untana 157 (CR, MO), Zona Prot. La Cangreja, vic. Quebrada Grande, ca. 2 km NNE of Mastatal de Puriscal, Grayun 8638 (CR, MO), ZP La Cangreja, ca. 1.5 km E of Santa Rosa de Puriscal, Grayun et al. 8336 (CR, MO), San Jose rd. from Parrita to Santiago de Purriscal, Barringer 1790B (CR), Res. Biol. Carara-Est. Quebrada Bonita, Chac6n 1406 (CR), Cordillera Talamanca, Rfo Hermoso, Finca E1 Quizarra, L. Willianes et al. 28479 (F, NY, US); Acosta, along Rfo Parritilla, ca. 1 km date at base; upper surface matte to weakly glossy, dark green, drying dark gray-green to sometimes blackened; lower surface matte, paler, drying yellowish gray-brown to dark yellow-brown; midrib 5- 8 mm diam., flat to broadly and obscurely sunken at base, weakly raised toward apex above, concolorous to slightly paler than surface, frequently much paler than surface or even white toward apex, E of Zoncuano, Grflyum el al. 11174 (INB, MO); Perez Zeled6n, Fila Tinanlaste, Valverde 741 (CR, MO); Puriscal, Fila Tufares, Salitrclles de Puriscal, Goncez-Laurito 7792 (CR); Z.P. La Cangreja, Cerros de Puriscal, San Martfn de Puriscal, La Fila Vala Blanca, J. Morales 2035 ((21t, INB, MO). Cultivated specimens. Costa Rica. 800-900 z1, cultivated at Munich as 1203/74, Horich s.n. (M). often with a light green streak distally! drying paler than surface or darker than surface al)ove, convex to bluntly round-raised, drying somewhat flattened with acute ribs below, drying darkel than surface below; primary lateral veins 7 to 12(15) per side, departing midrib at a 45°-55° angle (to 30°-50° at apex, 50°-90° at base), arising a( utely, then 14. Dieffenbachia isthlnia Croat, sp. nov. TYPE: Panama: along trail betw. Rfo Maje & Quebrada Brava, 60 m, 4 May 1976, 1: B. Croat 34656 (holotype, M0-240198!; isotypes, B!, K!, PMA!, US!). Figures 14, 28A. spreading weakly or not at all sunkell often greenish toward apex, sometimes raise(l 1leal midrib, and diminishing toward margins above; lllinor veins obscure above, obscurely visible to not visible below. INFLORESCENCES straight to sliglltly curved, 3 to 5 per axil, bracteoles 9-20 cm long; peduncle Planta 2-3(4.5) plerumque cm diam., ad 1 m, atroviridia; internodia petiolus 0.5-3 11-34(40) cm longa, cm lon- (2.5)4-12.5 cm long, 6-10 mm diall., sometimes flattened on one side in cross sec.ti(-)n, green; spathe gus, vaginatus 0.4-0.77 longitudinis, vagina 8.5-21 cm 15-17.5(23) cm long, medium green outside, slightlonga, lamina oblongo-ovata vel anguste ovata, (12)15- 30(39) cm longa, (6)10-26 cm lata, plerumque subcordata, nervis primariis lateralibus 7-12(15) utroque, inflorescentia 3-5 per quoque axillam; pedunculus (2.5)4ly paler green inside throughout, except the apical portion white at anthesis, to 1.7 c m lollger than the spadix, elongating somewhat after closing; spathe 12.5 cm longus, spatha 15-17.5(23) cm longa, spadix tube less than 2 cm diam. when furled, spathe 10.5-15.8 cm longus; pars feminea 6.5-9.5 cm longa, 1.4 blade 2.5-3 cm diam. at anthesis; s)
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia Figure 14. Dieffenbachia isthmia (Croat & Zhu 77115). A. Leaf blade adaxial surface. B. Close-up of petioles showing inflorescence. C. Close-up of female portion of spadix showing pistils and staminodes. D. Infructescences, one cut off to show the fruits. 719
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