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736 Annals of the Missouri Botanical Garden bands); sheath 6.7-10 cm long, the tip usually Cocle) to northern Colombia (Antioquia and Choco) rounded to inequilaterally emarginate, rarely de- from 30 to 800 m in Tropical rnoistforest (T-mf) and current, frequently with one side emarginate and Tropical wet forest (T-wf) life zones (Holdridge, with the other side acute; unsheathed portion lack- 1967). ing or to 5.2 cm long, sharply to bluntly sulcate; Phenology. The species has been seen in flowblades narrowly elliptic, 20-30 cm long (averaging er from May to December and fruiting collections 24 cm long), 5-8 cm wide (averaging 7.5 cm wide), have been reported from from September to May. 2.4-5.2 times longer than wide, 1.6-3.1 times lon- Discussion. Dieffenbachia obscurinervia is charger than petiole, slightly inequilateral, one side acterized by its scurfy brown internodes, conspic- 0.7-1 cm wi(ler than the other side, subcoriaceous, uously splotched petioles, and narrow blades with weakly bie olol ous, gradually acuminate, briefly apic- weakly developed primary lateral veins. It is not ulate at al)ex, {lequently somewhat falcate, cuneate easily confused with any other species but was into obtuse ol X ounded, often weakly inequilateral, correctly called D. pittieri, a species considered rerarely weakly sube ordate at base, upper surface dark stricted to the Isthmus of Panama, for many years. green, semit,lossy, drying dark yellow-green to dark See that species for a discussion of the differences. olive-greels; lowel surface slightly to moderately paler, drying yellow-green, midrib slightly raised to flat above, con( ololous, drying concolorous above, convex to narlowly rounded below, dark green, densely Paratypes. PANAMA. Canal Area: Pipeline Rd.7 mi. 1-37 N of GarYlboa7 S. Knapp 1042 (MO); Pipeline Rd. at Rio Agua Salud7 Croat 12352A (MO); Barro Colorado Island7 Armour 'I'rail 12807 Croat 8623 (MO); Drayton Trail pale yellowish green-spotted, drying paler than sur- 10007 Croat /2660 (MO); Drayton 19107 Croat 6775 (MO7 STRI); Draytoll 1490, Croat 5759 (M()). Cocle: vic. E1 face or datket than surface below; primary lateral Valle de AntI)ll, La Mesa7 Finca Maarenita7 Croat & Zhu veins 10 to 13 I)er side, weakly etched above, weak- 76666A (M ( )* I'MA). Colon: Santa Rita Ridge7 Croat ly raised or llot at all raised below; the interprimary 13849 (MO); Nombre de Dios7 Cro(lt t57.X29 (MO); Cocle veins alnsost as conspicuous as primary lateral del Norte7 ZI(lrnmel 4484 (MO); I'ollofelo-Nombre de Dios7 1.2 lzli. I)eyond jet. of rd. lo Isla Grande7 Croat veins, milsor veins moderately distinct below. IN- 49791 (M()7 I'MA); Rio Guanche7 (,. .S kln upstream from FLORES(E , N (E IiS 1 to 3 per axil; peduncle (1.7)3- rd. to Portobeloe Hammel & Train(r /4786 (MO); Loma 5 cm longe somewhat flattened in cross section; de la GloliaF near Fato (Nombre (le l)ios)7 Pittier 3847 spathe 11.5-2() c m long, 3.2-9.5 times longer than (US). Panalll.'l: Balboa West7 Za/l(lt(l (t (ll. 289 (PMA); peduncle, (lark green or pale green throughout, along trail ol'l' I,larlo-Carti Rd.7 4.6 rrli. from jet. with Pancaudate-ae lllninate, sometimes sharply reflexed below apex, .V)adix 10-18 cm long; free portion 5-9 cm long; pistillate portion 5-5.5 cm long; sterile Am. Hwy.7 M(l'l1(rson & Merello 8/4.S (MO). San Blas: Rio Cangandl (2angandi7 H. Herrer(l 245 (MO7 PMA); Nusigandi7 alotlg ll Llano-Carti Rd.7 ().7 km beyond Cuna headquarterst 11.6 km from Pan-Am. Hwy.7 Croat 75149 segment less than 1 cm long, 6 mm diam., with (KS MO). C()l,()MBIA. Antioquia: Slo Anorl valley7 near scattered staminodia throughout; pistils 20 to 40, moderately ( losely spaced, scattered, 2-2.6 mm diam.; stigma depressed-globose, 2 mm diam., staminodia white, usually 3 per pistil, 1.5-2(3.4) mm Planta Provi(lell( ia7 Shepherd 438 (M(), WIS); Anori7 Corr. Prov.7 Soejart(l & Renteria 3556 (HUA); %aragoza7 Rio Anori7 Quebra(ltl l, 'rirana-Rio Anori7 2 km N of Quebrada La Tirana7 3 k lll ll priver from Planta Providencia7 Alverson et al. 283 (C()l,); Carepa7 Tulenapa Reserva (ICA)7 Turbolong, flattene(l, oblong to clavate, grooved medially, Mutata7 40 kzl S of Turbo7 Callejas et al. 4873 (MO7 NY). Choco: Nu(ltlt, Quebrada Chaqui7 Galeano et al. 4825 drying flattened, thickened at apex; synandria ir- (MO); Quibddx, l, Concepcion7 15 km E of Quibdo7 Archer regularly rounded with margins undulate, minutely 2215 (US); Cor(loba7 Tierralta7 Rio Esmeraldas-Rio Simi7 warty, widely sunken at apex, 1.8-2.6 mm diam., 2 km above ( orlfluence7 Bernal et al. 1158 (COL). dark yellow-brown, matte on drying. INFRUC- TESCENCE broadly arching; spathe mostly decid- 20. Dieffenbachia oerstedii Schott, Oesterr. uous; spadix to 8 cm long; berries red, broadly el- Bot. Z. 8: 179. 1858. TYPE: Guatemala. Agualipsoid, to 1.5 cm long, less than 1 cm diam. cate, A. S. Oersted s.n. (holotype, C!). Figures Common name. Abior (Kuna) (de Nevers et al. 20, 29A. 7409). Herb, 30-75(100) cm tall (usually less than 50 Distribution and habitat. Dieffenbachia obscu- cm); stem erect or partially reclining, often conrinervia ranges from central Panama (as far west as spicuously clustered with numerous plants; sap Figure 20. Dieffenbachia oerstedii. A. Potted plant with inflorescences and leaves with mottled blades (Croat 68449). B. Potted flowering plant with pale midribs (Croat & Zhu 76794). C. Leaves and post-anthesis inflorescence showing ovate-lanceolate blades with scarcely sunken primary lateral veins (Croat 63208A).-D. Close-up of
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia Figure 20. (Continued) stems showing the free-ending petiole sheath (Croat 36297). EX F. (Croat 68449). E. Flowering plant with open inflorescences. F. Close-up of flowering plant showing open spathe blade and exposed stamlnate portlon of inflorescences. 737
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