l - CNCFlora

698 Annals of the
Missouri Botanical Garden
size (to 1.3 m vs. 0.75-1 m in D. oerstedii), and Raven 21532 (F), Rinc6n de Osa, Quebrada Apariciomore
coriaceous blades typically broadest at the
Quebrada Aguabuena, Gray7lm et al 3982 (CR, MO), vic
of Boscosa, at Quebrada Aguabuena, Croat & D. Hannon
middle (vs. broadest below the middle in D. oerste- 79238 (COL, INB, MO, NY, PMA, TEFH, TEX, UB), Secdii)
with more numerous primary lateral veins (6 to tor Esquinas, vic. of Fila Gamba hills behind Esquinas
12(14) vs. mostly 6 to 9 for D. oerstedii). In addi- Rain Forest Lodge, along Quebrada Negra, at end of side
tion, the blades of D. concinna are glossy above rd. off of Villa Bricena to Golfito Rd., Croat & D. Hannon
79291 (CM, DUKE, INB, MO, P), Refugio Nac. Golfito, S
and almost rounded at the base versus weakly tributary of Rio Canaza, Grayum et al. 9250 (MO), W side
glossy to matte and subcordate in D. oerseedti. Fila Gamba, ca. 6 km from Golfito airport, Croat & Gra-
Grayum et al. 10588, from the headwaters of the yalm 59930 (CM, K, MO), Rfo Claro, 2.5 mi. SE of Golfito,
Rio Piedras Blancas in Puntarenas Province, is un- Croat 67594 (MO), Orotina-Jaco, valley of Rio Grande
usual not only in occurring much higher (to 900 m)
Tarcoles, 1 km S of Quebrada Ganado, vic. Hotel Pink
Paradise, Croat 79076 (INB, MO), Par. Nac. Corcovado
than most collections of this species but in having Alonkey Woods, Kernan & Phillips 831 (CR, MO), Lower
a broader leaf (to 19 cm wide) with a longer petiole Lookout Trail, Kernan 748 (CR, MO); Est. Sirena, Que-
(to 30 cm). In other respects it agrees with other sada 51 (INB, MO, US, CR), Penfnsula de Osa, Estacion
collections of D. concinna. Hammel 9660, collected Biologica Los Patos, 8o34'N, 83o31'W, 200 m, 2 Sep.
1993, R. Aguilar 2195 (CR, INB, MO); 0-1 km W of park
at La Selva in Heredia Province in Cesta Rica, headquarters at Sirena, Liesner 2871 (CR, MO), Pan-Am
probably represents a hybrid between D. beachiana E4wy.! Rinc6n, Croat & D. Hannon 79167 (MO), Rinc6nand
D. concinna. The specimen has the slightly Rancho Quemado, just S of Rincon near Rio Rinc6n,
scabrous petioles and the impressed major veins of Croat & D. Hannon 79170 (HUA, INB, MEXU, MO,
D. beachiana, but the blades are more or less el-
QCA, RSA, WU), vic Palmar Norte, Allen 5669A (F, UC),
Punta Banco, M. Cha1a(lrrfa et al 257 (CR); Coto Brus,
liptic and dry more the color of D. concinna. Croat Guaymi Reserve, river trail along Rio Lim6ncito near jct.
& Hannon 79199, collected south of Palmar Norte, Of Villa Palacios school trail, Koshear 59 (CR), Golfito,
appears to be a hybrid between D aurantiaca and Refugio de Vida Silvestre, Marten 789 (CR), Carr. Inter-
D. concinna. Its blade shape and maculations, and am.-Sinai, Fila Huac.as! (a. 4 km NE (3Dy road) at Las
Huacas (;'Venecia"), (;relum & Evans 10156 (CR, MO);
numerous inflorescences suggest l). concinna, while Palmar Norte-3alisco, (Juebrada Benjamin, Graymm et al.
the broadly sulcate petiole favors D. aurantiaca. 9962 (CR, MO). San Jose: above Palmar Norte, Croat
35708 (MO). NICARA(tUA. Rio San Juan: Res. in{lio-
Additional specimens examined. COSTA RICA. He- Mafz, Cano el Tambor t)ranch of Rio San Juan, Rueda et
redia: near Puerto Viejo along rd. near Rfo Sucio, Croat al. 4070 (MO). PA N A MA. Panama: rd. to Puerto
35688 (MO), Croat 35702 (MO); Finca La Selva, OTS 3imenez, Osa, 40 km W of Panam. Hwy., rt. 2, L. Gomez
Field Station on the Rfo Puerto Viejo just E of its junction 19531 (MO).
with the Rfo Sarapiquf, Hammel 9688 (DUKE), Croat Cultivated collections. Costa Rica. Puntarenas: Esqui-
78732 (INB, MO, W). Limon: La Colombiana Farm, Unit- nas, 25 km SE of Palmar Sur, along Pan-Am Hwy., origed
Fruit Co., Standley 36739 (US); headwaters of Quebra- inally collected by Bru( e. McAlpin (Selby 85-33), 25 m,
da Mata de Limon, Finca Anai (Sixaola region), Grayum 2 Jan. 1994, cultivate(l at MBG, vouchered as Croal
et al. 4447 (MO); Cerro Tortuguero junto a la Barra de 77281 (MO); 15 May 1996, Croal 78318 (MO).
Tortuguero, R. Robles 2090 (CR, INB, MO), Barringer et
al. 1973 (F); Par. Nac. Tortuguero, Go'mez-Laurito 7856 5. Dieffenbachia copensis Croat, sp. nov. TYPE:
(CR); Boca de las Lagunas de Tortuguero, Burger & An-
Panama. Cocle: E1 Cope, forest on Continental
tonio 11249 (F, MO); Pococi, Refugio de Vida Silvestre
Barra del Colorado, SE base Of Cerro del Tortuguero, Gra- Divide above El Cope, 8°38'NX 80°38'W, 700yum
et al. 11139 (CM, CR, INX, MO, USJ), Rfo Chirri- 900 m, 27-29 Apr. 1985 B. Hantmel 13636
p6cito-Rfo Sardina, Refugio Nac. Barra del Colorado, Gra- (holotype, M0-3284876!). Figures 6, 27B.
yum 9777 (CR, MO); Rio Reventaz6n, Finca Montecristo
below Cairo, Standley & Valerio 48960 (US). Puntaren- Herba 45 cm alta; internodia 7 mm longa, 7 mm diam.,
as: McAlpin 85-33 (SEL); 2 km NW of Chacarita, 30 km petiolus (17)21-29.5 cm longus, vagina S12.5 cm longa;
S of Palmar Sur, Grayum & Fleming 8119 (CR, INB, MO); pars libera 10.5-15.5 cm longa, lamina anguste oblanceoridge
betw. Rio Riyito (valley of Laguna Chocuaco) & lata, 25.5-33.3 cm longa, 5.2 cm lata, nervis primariis
Quebrada Banegas, S of Cerro Rancho Quemado, Grayum lateralibus 9-12 utroque, inflorescenlia 1 per asillam, peet
al. 7567 (MO); Rinc6n de Osa Rancho Quemado, ca. dunculus 5 cm longus, spatha 15.7 cm longa, 1.2 cm lala;
10 km W of main Rincdn-Pto. Jimenez Rd., Croat & Gra- spadix 12.3 cm longus.
yum 59785 (CR, MO); Osa, 6 km W of Rinc6n, Grant &
Rundell 92-01928 (CR, MO, US); near the airfield ca. 4
Small herb, to 45 cm tall; internodes about as
mi. W of Rinc6n de Osa, Burger & Stolze 5461 (CR, F), long as broad, 7 mm diam., drying matte, dark
Osa Penfnsula, NW of airfield, ca. 5 km W Of Rinc6n de brown, finely and irregularly folded; petioles
Osa, Burger & Liesner 7196 (CR, F, MO, PMA); Osa Pen- (17)21-29.5 cm long, sheathed 41Wo-56Wo their
insula, Nicolson 3393 (BM, US); Costena-Cruces, Rfo Pie- length, sheath 8-12.5 cm long, narrowly acute at
dras Blancas, Cerro Anguciana, Grayum et al. 10588
(INB, MO, CR); Rio Sandalo, Dodge 10198 (F, MO), apex, drying dark brown to medium yellow-brown;
Dodge 10020 (F, MO); Palmar Norle de Osa, Allen 5669 free portion 10.5-15.5 cm long, subterete, obtusely
(EAP, UC, US); Osa Peninsula, 4 mi. W of Rincon de Osa,
sulcate, drying medium yellow-brown to dark