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726 Annals of the Missouri Botanical Garden Holm-Nielsen et al. 2769 (AAU, NY, Q(:NE, S)7 Hacienda wide), 1-2 times longer than petioles, coriaceous to Monica, 12 km E of San Carlos, Sparre 19397 (S); Haci- subcoriaceous, semiglossy, bicolorous, weakly inenda Clementina, Harling 313 (S); Babahoyo-Montalve, Sparre 14556 (S), Cornejo & Bonifaz 4828 (GUAY, MO); equilateral, one side 0.6-3.5 cm wider than the oth- Centro Cientifico Rio Palenque, Croat 38669 (MO, er, usually short-acuminate at apex, sometimes QCNE), Dodso7l & Tan 5338 (SEL), Croat 73826 (MO, acute to rounded with a short acumen, acute to QCNE), Croat 50655 (MO), Fallen & Ray 860 (SEL); rounded at base, with the edges turned up near the Quevedo, Asplund 15574 (S)7 Dodson 6188 (RPSC, SEL); Vinces, Mocachi-Palenque, Jauneche forest, km 70 Quevbase; upper surface dark green, drying gray-green edo-Palenque, via Mocachi, Dodson et al 10594 (GUAY, to dark olive-green, rarely yellowish brown; lower MO, SFL). Manabi: Portoviejo-Pichincha rd., ca. 20 km surface slightly paler, drying yellowish green, rarely E of San Placido, Harling & Andersson 24778 (GB, MO); yellowish brown7 midrib flat, 1-2 cm wide at base, Cerro Montecristo, S of Manta, Sparre 19488 (S); Mach- concolorous or slightly paler than surface above, alilla NP, zona de San Sebastian, Gentry et al. 72496 (MO, QCNE); Chone-Santo Domingo Rd., ca. 20 km NNE of drying slightly paler than surface and weakly raised Flavio Alfaro, Montanas de Convento, Harling & Anders- above, concave to prominently raised on lower surson 18898 (GB); Hacienda Don Juan, 10 km NE of Jama, N of Rio Don Juan, Neill et al 11683 (QCNE); 223 km S of Pedernales, 3.5 km SW of Camarones, I)elinlss 452 (AAU, NY, MO). Napo: Auca, Plowman 14121 (F). Pastaza: Coca-Rio Tiguino, t35.23 km S of Coca, Croat 72573 (MO). Pichincha: vic. Hotel Tinalandia, 9.6 km E of Santo Domingo de los Colorados, Croat 55666 (GUAY, MO, QCA); Machachi-Santo Domingo, 19.3 km E of Alluriquin, Thompson & Rawlins 1104 (MO); Centinela, Montanas de Ila, 12 km E of Patricia Pilar, L0jtnant & Molau 15839 (AAU); Santo Domingo-Quevedo, Patricia Pilar, face, drying brownish; primary lateral veins 15 to 26 pairs, sunken above, convex below, arisin; at an acute angle to the midrib then spreading at an angle of 45°-60°(70°), sometimes to 90° near the base of the blade, sometimes drying moderately wrinkled; interprimary veins lacking or 1 between each pair of primary lateral veins, sometimes almost as prominent as the primary lateral veinsS minor veins indistinct Juvenile telades with acute base and sol- Dodson et al. 14638 (M0, QCNE), Gentry et al. 26709 id green midrib. INFI,ORESCENCES 1 to 3 per (MO); vic. of Santo Domingo de Los Colorados, Peripa, axil; peduncles (5.5)7-25 x 1.5-2 cm; spathe SW of Santo Domingo, Croat & Nunez 82064 (MO, QCNE); Nanegalito-Pto. Quito Rd. km 113, ENDESA, 5 medium to dark greerl broadly curved, long-acukm W of San Vincente Andoas, Croat 82829 (MO), Croat minate, 2748 cm long, 2.5-8.2 times longer than et al. 83795 (MO); Rfo Silanche, Quito-Pto. Quito, km peduncle, to 4 cm wi({e at anthesis, tube flattenin; 113, Rodriguez 262 (NY, QCA); Santo Domingo-Quininde 5-12.5 cm wide, oorlstlicted area 2.5-3 cm diam., rd., km 41, Zak et al. 5414 (QCA); vic. Maquipuquna flattening 3.2-5 ( m wide, spathe blade 3-6 cm Res., rd. to Maquipucuna Lodge, LSeimbeck R. 306 (AAU). Sucumbios: Lago Agrio-Baeza, ca. km 107, Croat 50480 wide at anthesis, flattening to 6-12 cm wide mid- (MO). VENEZUELA. Maiquetia, Andre 457 (K). way, the distal inner surface sometimes white when Cultivated specimens. Ecuador. Napo: Auca Oil Field, open; spadix (21)35-38 cm long; free portion 12- 23 Sep. 1991, Ingram 1124 (MO). 19 cm long; pistillate portion of spadix (except sometimes the uppermost part) fused to spathe 13- 16. Dieffenbachia longispatha Engl. & K. 15 cm long; fertile s;taminate portion (8)11-14 cm Krause, Pflanzenr. IV, 23 Dc(Heft 64): 44. x (9)12-14 mm ((3rying 6-9 mm diam.); mostly 1915. TYPE: Panama. Colon: Fato (Nombre de sterile intermediate portion (2)3-4.3 cm long with Dios), July 1911, H. F: Pittier 3838 (holotype, a few scattered staminodia in the upper half (some- US!; isotypes, B!, F!, M0!). Figures 16, 29B. times to throughout ite; length); pistillate flowers 10 Terrestrial, (1)1.5-3.5 m tall; sap very foul and to 26, round or barely bilobed, widely spaced 5- pungent; stem prostrate at base, then erect; inter- 10(20) mm apart, forming in a single irregular row nodes 4-12 cm diam., with leaf scars prominent, or scattered but usually no more than 2 flowers dark green, semiglossy, drying dark brown to or- across the width of the spadix (rarely 3); ovaries ange-brown; petioles thick and succulent, semiglos- pale green, 4-7 mm diam.; stigmas 4-6 mm diam., sy, usually solid dark green, rarely streaked with yellow to orange, somewhat broadly bowl-shaped, pale green, 23-55 cm long (averaging 36 cm long), 5-7 mm thick on the edge, medially with 1 to 2 sheathed to about middle (0.58-0.85 their length, somewhat elongate lobes, the lobes 1-1.5 mm averaging 0.72); sheath 25X1 cm long (averaging diam., somewhat longer than broad; staminodia 5 25 cm long), inequilaterally rounded at apex, some- to 6 per pistil, white, irregular, 2-6 x 2-3 mm, times weakly free-ending; unsheathed portion 4.0- much flattened at base, less so toward the apex, 30.5 cm long (averaging about 11 cm), C-shaped often somewhat puckered at the apex; synandria in and obtusely sulcate or + terete with a faint flat spirals of 4 to 7 each, 3-4 mm wide, subrounded, rib adaxially; blades oblong-elliptic, 41-72 x 17- drying light yellow-brown and concave at apex. IN- 38 cm (averaging 53 x 24 cm), 1.7-2.9 times lon- F1lUCTESCENCES 17-24 cm long; berries 1.5-2 ger than wide (averaging 2.3 times longer than cm diam., often deeply emarginate at both ends and
Volume 91, Number 4 2004 Croat Revision of Dieffenbachia Figure 16. Diedfenbachia longispatha. A. Habit with open inflorescence. B. Plant habit with cluster of inflorescences. C. Crown of plant with close-up of leaf base with open inflorescence. D. Inflorescence at anthesis with staminate spadix exposed. A, C. (Croat & Zhu 76257); B, D. (Croat & Zhu 76203). 727
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