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A B S T R A C T B O O K – A B S T R A C T S O F T A L K S INTEGRATED MOLECULAR CIRCUITS FOR STEM CELL ACTIVITY IN ARABIDOPSIS ROOTS Ben Scheres University of Utrecht, Utrecht, The Netherlands E-mail: b.scheres@uu.nl 25 X X I V S P P S C O N G R E S S 2 0 1 1 Pleanary session 3 Plant stem cells reside in niches and are maintained by short-range signals emanating from organizing centres. The Arabidopsis PLETHORA genes encode transcription factors required for root stem cell specification [1, 2]. PLT expression is induced by the indolic hormone auxin, depends on auxin response factors and follows auxin accumulation patterns. The PLT gene clade extensively regulates expression of the PIN facilitators of polar auxin transport in the root and this contributes to a specific auxin transport route that maintains stem cells at the appropriate position[3]. We are addressing the properties of the PLT-PIN feedback loop by gene and protein network analysis and computational modelling. The emerging picture is one in which flexible feedback circuits translate auxin accumulation into region- and cell type specification patterns. Stem cells and their daughters in the root display specific asymmetric divisions at fixed locations. We investigate how such divisions are spatially regulated. The SHORTROOT- SCARECROW transcription factor pathway plays a role in patterning the quiescent center and cortex/endodermis stem cells and provides mitotic potential to the stem cell daughters that form the proximal meristem. This activity involves the conserved RETINOBLASTOMA-RELATED (RBR) pocket protein[4], and we have established molecular links between the RBR pathway and SCARECROW action that form a feedback control system. In addition, RBR activity is modulated by auxin abundance, itself regulated through an intercellular distribution system, and by cell cycle progression. Formal analysis of this feedback circuit indicates that it acts as a bistable switch that ensures the occurrence of an asymmetric division at fixed positions. Our work illustrates how formative divisions that shape plant tissues can be robustly positioned by dynamic regulatory circuits that combine intracellular and extracellular loops. References [1]Aida et al.,Cell 2004 [2] Galinhaet al., Nature 2007 [3] Grieneisen et al., Nature2007 [4] Wildwater et al., Cell2005
Pleanary session 4 A B S T R A C T B O O K – A B S T R A C T S O F T A L K S CHLOROPLAST PROTEIN IMPORT Jürgen Soll Ludwig-Maximilinas University, München, Germany E-mail: soll@lmu.de Chloroplasts must import several thousand different proteins to obtain and maintain their biochemical identity and function. Protein turnover, environmental adaptation and cellular division of labour require a constant adjustment of the protein complement in chloroplasts. Protein import into chloroplasts is regulated at all stages of the process, ranging from co- or early posttranslational events in the cytosol to late stages in the translocation process which reflect the metabolic as well as the redox status of the organelle. 26 X X I V S P P S C O N G R E S S 2 0 1 1
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Page 3 and 4: Foreword eword ord A B S T R A C T
Page 5 and 6: Program of the Congress gram of the
Page 7 and 8: A B S T R A C T B O O K - P R O G R
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Page 11 and 12: Plenary session 1 A B S T R A C T B
Page 13 and 14: Plenary session 2 THE BIOLOGY OF AL
Page 15 and 16: Session 1. Biotechnology and synthe
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Page 19 and 20: Session 2. Systems biology and -omi
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Page 29 and 30: Invited talk 4 A B S T R A C T B O
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Page 37 and 38: Session 7.Development A B S T R A C
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Page 41 and 42: Pleanary session 6 A B S T R A C T
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SCF = A B S T R A C T B O O K - A B
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A B S T R A C T B O O K - L I S T O
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Sponsors onsors ors A B S T R A C T
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