382 400 mm SL. Large specimens are recorded of C. orinocensis (617 mm by Machado-Allison, 1971; 410-565 mm by Jepsen et al., 1999), and C. intermedia (429 mm by Jepsen et al., 1999). Cichla intermedia Machado-Allison, 1971 (Figs. 90-92) Cichla intermedia Machado-Allison, 1971: 473, fig. 7 (río Casiquiare, Territorio Federal Amazonas. Holotype MBUCV V-6489). Material examined. Six specimens, 87.6-192 mm SL. Venezuela: Rio Orinoco drainage: MBUCV-V 3477, paratypes, 3, 87.6-137.3 mm SL; Apure, Rio Cinaruco; 6 Mar 1966, V. E. Ogilvie. – MBUCV-V 6498, 1, 192 mm SL; Apure, Rio Cinaruco; 26 Feb 1964. C. Rivero. – MBUCV-V 6496, paratype, 1, 153 mm SL; Amazonas, Quiritare, 4 h before mouth of Rio Cunucuna; 3 Oct 1966, R. Campos & C. Duque. – MBUCV-V 12819, 1, 88.9 mm SL; Amazonas, Rio Cataniapo, ca 300 m from mouth, in small caño. 3 Jul 1981, R. Royero et al. Diagnosis. Juveniles with uninterrupted dark band along middle of side, young and adults uniquely distinguished among species of Cichla by colour pattern with lateral band transformed to row of irregular dark blotches and six or seven narrow dark vertical bars across side below dorsal fin (corresponding in position to bars 1, 1a, 2, 2a, 3, additional bar below end of soft dorsal in base, and occasionally one anteriorly on caudal peduncle). Adults with several small black blotches posterior to orbit and on gill cover. Light spots on side absent at all sizes (vs. present in most other species of Cichla). Lateral line almost always continuous; scales in E1 row 96-108. Description. Refer to Figs. 90-92 for general shape and colour pattern, Table 26 for morphometrics, and Tables 2-10 for meristics. Relatively elongate (depth 25.7-30.7 % SL, N = 6, 88-192 mm SL). Frontal contour straight ascending, gently curved between eye and dorsal fin origin. Nuchal hump absent in specimens observed by us, but present in specimens on published photographs (Román, 1981: 68). Maxilla reaching to below middle of orbit, lower jaw to below posterior margin of orbit. Lateral line continuous on both sides in four specimens, continuous on one side in one specimen, and bilaterally discontinuous in one. Dorsal-fin spine 4 longest; soft dorsal fin rounded, reaching about middle of caudal peduncle in small specimens, slightly beyond middle in specimen 192 mm SL. Soft anal fin rounded, extending as far as soft dorsal fin. Caudal fin emarginate, with angled corners in specimen 89 mm SL, concave with angled upper corner and rounded lower corner in 137 mm specimen, in larger specimens generally injured, shape not unambiguously determinable. Pectoral fin rounded in specimen 89 mm SL, rays 5 and 6 longest, reaching about halfway to anterior anal fin rays; in larger specimens subacuminate or acuminate, ray 4 longest, reaching halfway to soft anal fin. Pelvic fin rounded in specimen 89 mm SL, rays 1-2 longest, reaching about halfway to anterior anal fin rays; in larger specimens becoming subacuminate, tip terminated by ray 1, extending Table 26. Morphometry of Cichla intermedia. Measurements are in percent of SL, except SL (in mm). Linear regression parameters calculated from original measurements in mm. N min max mean SD a b r SL (mm) 6 87.6 191.9 125.2 Head length 6 31.5 35.5 33.0 1.51 2.7221 0.3058 0.994 Snout length 6 11.0 12.3 11.5 0.55 –0.2685 0.1175 0.992 Head depth 6 20.5 23.2 21.7 1.19 –3.3703 0.2461 0.995 Body depth 6 25.7 30.7 27.2 2.05 –5.3279 0.319 0.988 Orbit diameter 6 7.8 10.0 9.0 0.89 3.2846 0.061 0.999 Interorbital width 6 6.5 8.2 7.3 0.64 –2.3048 0.0932 0.997 Pectoral fin length 6 17.2 23.6 20.5 2.21 –7.7116 0.2727 0.997 Upper jaw length 6 14.1 15.5 14.6 0.57 –0.4633 0.1504 0.995 Lower jaw length 6 18.2 20.0 18.8 0.66 0.1854 0.1859 0.997 Caudal peduncle depth 6 9.1 10.9 10.2 0.65 –1.8005 0.1178 0.996 Caudal peduncle length 6 16.8 19.5 18.2 1.16 4.0503 0.1464 0.997 Dorsal spine length 6 11.1 12.9 11.9 0.89 –1.8308 0.1351 0.987 <strong>Kullander</strong> & Ferreira: Review of Cichla
Fig. 90. Cichla intermedia, MBUCV-V 12819, 88.9 mm SL; Venezuela: Amazonas: Rio Cataniapo. Fig. 91. Cichla intermedia, paratype, MBUCV-V 3477, 137.3 mm SL; Venezuela: Apure: Rio Cinaruco. Fig. 92. Cichla intermedia, MBUCV-V 6498, young female, 192 mm SL; Venezuela: Apure: Rio Cinaruco. about halfway to anal fin origin. Scales absent from dorsal fin in small specimens, in specimen 137 mm SL a few scales present basally on soft dorsal fin, in specimen 192 mm SL double rows in single band behind each soft ray, Ichthyol. Explor. Freshwaters, Vol. 17, No. 4 383 except on last four interrradial membranes, squamation extending for about ‚ of length of rays. Scales present on anal fin in all specimens, in 89 mm specimen only on spinous portion and bases of anterior soft rays, in specimen 192 mm
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Verlag Dr. Friedrich Pfeil ISSN 093
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Ichthyol. Explor. Freshwaters, Vol.
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that of Ringuelet et al. (1967) cle
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except in C. intermedia. Occipital
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Prominent rostral fold on maxilla (
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Principal Component 2 2 1 0 -1 -2 -
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Frequency 50 40 30 20 10 0 60 70 80
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soft dorsal fin; bars 1 and 2 divid
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Fig. 7. Cichla ocellaris, NRM 28389
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10° 0° -10° -20° -30° -80° Ci
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uito, Rio Aguaro on the way towards
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Fig. 13. Cichla orinocensis, MCNG 2
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confluent elongate yellow or white
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Fig. 19. Cichla orinocensis, MNHN A
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Cichla nigromaculata Jardine, 1843
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10° 0° -10° -20° -30° -80° Ci
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of the principal vertical bars in C
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Fig. 26. Cichla monoculus, freshly
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on back, nape and dorsally on cauda
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Diagnosis. Distinguished from all o
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Several light spots on proximal hal
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Fig. 30. Cichla kelberi, NRM 42401,
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- Page 57 and 58: Fig. 47. Cichla melaniae, not prese
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- Page 61 and 62: Fig. 52. Cichla piquiti, paratype,
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- Page 67 and 68: Fig. 57. Cichla thyrorus, paratype,
- Page 69 and 70: Cichla jariina, new species (Figs.
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- Page 73 and 74: idge. - BMNH 1902.11.7:15, 1, 197 m
- Page 75 and 76: Fig. 67. Cichla pinima, paratype, N
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- Page 85 and 86: Fig. 78. Cichla vazzoleri, holotype
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- Page 89 and 90: Fig. 81. Cichla temensis, NRM 22921
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