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Fen Management Handbook - Scottish Natural Heritage

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grazing on vegetation composition and structure, and on molluscs.<br />

Measures of vegetation structure and composition were recorded in 20 quadrats<br />

in each of seven randomly located 15 x 30 m plots. One half of each plot was open<br />

to grazing, and the other half fenced off from grazing. Densities of molluscs were<br />

estimated by carrying out timed searches in eight 0.5 x 0.5 m quadrats in the grazed<br />

and ungrazed halves of each of these seven plots.<br />

The effects of grazing<br />

Statistical significant differences in the composition and structure of S25 tall-herb<br />

fen after four years of grazing were:<br />

– Increased dominance of reed sweet-grass Glyceria maxima at the expense of<br />

reed Phragmites australis.<br />

– Increased plant species-richness.<br />

– Reduced vegetation height<br />

– Reduced biomass of common reed and total vegetation biomass.<br />

– Reduced inflorescence density of common reed.<br />

Cattle were frequently observed feeding on the bark of willow scrub in winter,<br />

breaking willow branches with their horns and browsing their leaves, and so<br />

probably also reduced the growth of low scrub. No differences were detected in<br />

fine-scale (tens of centimetre) variation in habitat structure between grazed and<br />

ungrazed areas. However, the large-scale variation in grazing pressure within the<br />

fen increased large-scale (tens of metres) variation in habitat structure.<br />

Plant species showing significant differences in frequency between grazed<br />

and ungrazed plots after four years of grazing.<br />

Species more frequent in grazed<br />

halves of plots<br />

Celery-leaved buttercup Greater pond-sedge<br />

Common duckweed<br />

Gypsywort<br />

Pink/blue water-speedwell<br />

Purple loosestrife<br />

Reed sweet-grass<br />

Tufted forget-me-not<br />

132<br />

Species more frequent in ungrazed halves<br />

of plots<br />

Grazing substantially decreased densities of Desmoulin’s whorl snail. This species,<br />

though, remained widely distributed in areas within the grazing unit that the cattle<br />

did not graze.<br />

Conclusions<br />

Light grazing by Highland cattle has proved a useful method for increasing the plant<br />

species-richness and probably large-scale vegetation structure of derelict fen. It is<br />

probably also likely to slow the rate of succession to scrub.<br />

On the negative side, grazing reduced overall densities of at least some key mollusc<br />

species. These population reductions, though, have to be set against losses which<br />

would take place if the fen was left unmanaged to succeed to scrub. They also<br />

have to be set against likely increases in abundance of other important invertebrate<br />

species which benefit from more varied habitat structure. Breeding densities and

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