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Growth model of the reared sea urchin Paracentrotus ... - SciViews

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individuals related to <strong>the</strong> presence <strong>of</strong> o<strong>the</strong>r virtual individuals (inhibitorsinhibited<br />

interactions). With current <strong>model</strong> (eq. 31) and quantile regression<br />

method (eqs. 21 and 22), it is only possible to fit one curve at a time. An<br />

extension or adaptation <strong>of</strong> both <strong>the</strong> <strong>model</strong> and <strong>the</strong> regression method are<br />

required to link curves for different quantiles.<br />

In <strong>the</strong> case <strong>of</strong> parameter l, we have already mentioned that we expect<br />

l = 0 for τ = 1, according to <strong>the</strong> hypo<strong>the</strong>sis that larger animals in <strong>the</strong> batch<br />

are not inhibited at all (see above, construction <strong>of</strong> <strong>the</strong> <strong>model</strong>). We would<br />

also expect a monotonous increase <strong>of</strong> l with a decrease <strong>of</strong> τ because<br />

fractions <strong>of</strong> smaller individuals should be more inhibited than fractions <strong>of</strong><br />

larger ones (recall this is a size-based competition mechanism). Fig. 32A<br />

highlights a linear relationship between l and τ, except for <strong>the</strong> 10% smaller<br />

fraction. Consequently, we constrain l(τ) as:<br />

where s is <strong>the</strong> slope <strong>of</strong> <strong>the</strong> linear relation.<br />

Part IV: A growth <strong>model</strong> with intraspecific competition<br />

l( τ ) = s⋅(1<br />

− τ )<br />

(32)<br />

k1 and k2 appear negatively correlated in Fig. 32B but are quite<br />

constant along τ values, except for extreme quantiles. Moreover, large<br />

values <strong>of</strong> k2 for <strong>the</strong> three rightmost points in <strong>the</strong> graph at Fig. 32B seem<br />

associated with a potential overestimation <strong>of</strong> corresponding l values in<br />

Fig. 28A (outliers). In regard with <strong>the</strong>se considerations, reasonable<br />

relationships between k1/k2 and τ could be:<br />

k1( τ ) = cste = k1; k2( τ ) = cste = k2<br />

(33)<br />

with k1 probably different (and lower) than k2.<br />

158

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