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ASCOMYCETES<br />

ASCOMYCETES<br />

The Ascomycetes comprise a vast number of species of fungi that differ<br />

fundamentally from the Basidiomycetes in the manner in which the spores are<br />

formed. In the Basidiomycetes the spores are formed outside the mother cell or<br />

basidium, usually developing on Httle stalks that arise from it, whereas in the<br />

Ascomycetes the spores are produced within the mother cell or ascus and are<br />

not discharged until they are mature.<br />

The great majority of the Ascomycetes are minute fungi requiring a<br />

microscope for determination of their characters but some are large enough to<br />

attract the attention of the amateur collector and a few are known to be among<br />

our best edible fungi. Although a microscope is needed to actually see the<br />

difference between an ascus and a basidium, in practice it is not difficult to<br />

recognize an ascomycete in the field.<br />

All of the species described here, with one exception, belong in the section<br />

Discomycetes in which the asci are borne in an exposed fruiting layer or<br />

hymenium and not within a closed fruiting body. Relatively few species are<br />

described here and anyone interested in the group should consult special<br />

works deahng with them. The single pyrenomycete described, Hypomyces<br />

lactifluorum (Schw.) TuL, is not included in the key.<br />

Key<br />

1. Fruiting body with a distinct stipe and differentiated pileus 2<br />

1. Fruiting body more or less cup-shaped, without a differentiated<br />

pileus but sometimes stipitate 8<br />

2. Pileus pitted or honeycombed 3<br />

2. Pileus convoluted, wrinkled, or smooth, not pitted 5<br />

3. Base of pileus attached to stipe 4<br />

3. Base of pileus free from the stipe Verpa bohemica<br />

4. Pileus subglobose to ovoid; pits irregular with edges<br />

the same color or paler Morchella esculenta<br />

4. Pileus conical; pits more or less longitudinally<br />

arranged with edges darker Morchella angusticeps<br />

5. Pileus irregular, reddish brown, surface convoluted;<br />

growing on the ground in spring Gyromitra esculenta<br />

5. Pileus slightly wrinkled or smooth, usually more<br />

or less saddle-shaped 6<br />

6. Stipe deeply longitudinally fluted Helvella crispa<br />

6. Stipe smooth or slightly furrowed at base 7<br />

7. Pileus smoky gray to smoky yellowish or nearly black Helvella elastica<br />

7. Pileus tan to reddish brown Gyromitra infula<br />

8. Cups bright scarlet, whitish-hairy externally;<br />

growing on sticks in the spring Sarcoscypha coccinea<br />

8. Cups not scarlet 9<br />

9. Cups black, stipitate, tough Urnula craterium<br />

9. Cup brownish, sessile to substipitate, soft, fleshy,<br />

spreading out widely; growing on wood Peziza repanda<br />

257

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