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Medicinal Plants Classification Biosynthesis and ... - Index of

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Ethnobotany <strong>and</strong> Experimental Pharmacology <strong>of</strong> Echinodorus Gr<strong>and</strong>iflorus<br />

impeding the invasion <strong>of</strong> the air cavities <strong>of</strong> the aerenchyma for the water. The soluble<br />

fraction would be already related to the defense against herbivory in the submerged portion<br />

<strong>of</strong> the petioles.<br />

Chemical Constituents<br />

They were identified in the leaves <strong>of</strong> E. gr<strong>and</strong>iflorus:<br />

1) FATTY ACIDS: linolenic acid <strong>and</strong> dodecanoic acid (TANAKA, 2000); palmitic acid<br />

(PIMENTA, 2002).<br />

2) TERPENOIDS<br />

Essential oils: linalool, dihydroedulan, trans-caryophyllene, alpha humulene, Efarnesene,<br />

beta selinene, alpha farnesene, delta cadinene, E-nerolidol, caryophyllene<br />

oxide, humulene epoxide, bisabolone, drimenol, neocembrene, echinoic acid,<br />

cembranoid, phytol (PIMENTA et al., 2006).<br />

Diterpenoids: echinodol (MANS <strong>and</strong> HARTMANNS, 1993), echinoic acid<br />

(TANAKA et al., 1997), phytol, hardwikiic acid, (-) 15-etoxicleroda 3 acid, 13-dien<br />

- 15,16-olide-18-óic, acid (-) - (16)-hidróxi-cleroda-3,13-dien-16,15-olide-18 - oic<br />

(COSTA et al., 1999, TANAKA, 2000), acid (-) -15 hidroxicleroda-3,13 - dien-<br />

16,15-olide-18-oic, acid (-) -cleroda-3,13(16),14-trien-18-oic (TANAKA, 2000),<br />

chapecoderins A, B <strong>and</strong> C (KOBAYASHI et al., 2000c), echinodolides A <strong>and</strong> B<br />

(SHIGEMORI et al., 2002), solidagolactona-I (PIMENTA, 2002).<br />

3) STEROIDS: 24-etilcolest-4-en-3,6-dion; 3ß-the-ß-D-glicopiranosil sitosterol<br />

(TANAKA, 2000); campesterol, stigmasterol, sitosterol (PIMENTA, 2002).<br />

4) FENOLIC ACIDS: acid caffeic, acid ferulic <strong>and</strong> acid is<strong>of</strong>erulic (PIMENTA, 2002).<br />

5) FLAVONOIDS: isoorientin, swertisin <strong>and</strong> isovitexin (PIMENTA, 2002;<br />

SCHNITZLER et al., 2004), swertiajaponin, <strong>and</strong> isoorientin-7,3'-dimetil-ether<br />

(SCHNITZLER et al., 2004).<br />

6) ALKALOIDS: echin<strong>of</strong>illins A, B, C, D, F (KOBAYASHI et al., 2000°;<br />

KOBAYASHI et al., 2000b).<br />

7) DERIVED OF THE TARTARIC ACID: acids caftaric, chicoric, cafeoil tartaric<br />

feruloil, 2-the-feruloil tartaric <strong>and</strong> tartaric di-feruloil (SCHNITZLER et al., 2004).<br />

Ethnopharmacology<br />

The parts <strong>of</strong> E. gr<strong>and</strong>iflorus used for the treatment <strong>of</strong> several illnesses they are the leaves<br />

<strong>and</strong> the rhizomes. The use <strong>of</strong> the leaves is mentioned in FARMACOPÉIA DOS ESTADOS<br />

UNIDOS DO BRASIL (1926, 1959); in Coimbra (1994), Corrêa Júnior et al. (1994), Teske<br />

<strong>and</strong> Trentini (1995), Nogueira (2000), Almança <strong>and</strong> Carvalho (2003). The use <strong>of</strong> the<br />

rhizomes was mentioned by Corrêa (1984), Lorenzi <strong>and</strong> Matos (2002) <strong>and</strong> Dutra et al.<br />

(2006).<br />

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