Horticulture Principles and Practices

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Scale Compressed stem (a) Whole bulb (b) Cross section (c) Whole scaly bulb FIGURE 3–10 A scaly bulb. (Source: George Acquaah) FIGURE 3–11 Tubers. These swollen stems vary in size and shape depending on the species. The flesh color is also variable and may be creamish, whitish, yellowish, or some other color. (Source: George Acquaah) (c) Whole tuber of potato (a) Tuber of Solanum tuberosum (b) Tuber of Dioscorea spp. 7. Pseudobulbs. Also called false bulbs, these modified stems are commonly produced by orchids and are variable in appearance. The Dendrobium orchid produces an elongated and jointed structure with offshoots at the upper nodes. Once roots form at the base of these offshoots, they may be removed and transplanted. The most popular orchids (e.g., Cattleya, Miltonia, Laelia) can be propagated by using rhizomes or stolons divided into sections containing about five pseudobulbs. 8. Tubers. In plants such as yam (Dioscorea spp.) and white (or Irish) potato (Solanum tuberosum), the underground stems are highly enlarged as storage organs (Figure 3–11). Caladium is a popular tuber plant. Whereas Irish potato tubers are generally small in size, yam tubers vary widely in size and shape, according to the species, some attaining lengths of several feet. Tubers have nodes and axillary buds called eyes. Tubers develop best under reduced photoperiod (short days) and lower night temperatures. However, shoot development is vigorous under long-day conditions. 9. Tuberous roots. They lack nodes and axillary buds, the buds are present only at the shoot (proximal) end. 3.4 LEAF There are five types of leaves: foliage leaves, bud scales, floral bracts, sepals, and cotyledons. 3.4.1 FUNCTIONS The major functions of leaves are the following: 1. Food manufacture. The most widely known function of the leaf is photosynthesis. This function is performed by foliage leaves, which are the most readily visible type of leaf. 3.4 Leaf 75
2. Protection. Buds (vegetative or floral) are delicate tissues that need protection while developing. Several nonfoliage leaves assume the important role of protecting developing buds: a. Bud scales. Also called cataphylls, bud scales are modified leaves that protect buds (apical or axillary) while in the resting stage (dormancy) or vegetative buds. b. Floral bracts. Floral bracts, also called hypsophylls, protect the inflorescence during development. They perform a role similar to that of cataphylls. c. Sepals. Collectively called calyx, sepals operate like hypsophylls. Sepals are not durable, however, and often either senesce or abscise after the flower has fully expanded to its mature size. In certain flowers, sepals may be brightly colored like petals and may exude fragrances and produce nectar. 3. Storage. Storage as a function of the leaf is often overlooked. Cotyledons (or seed leaves) store food that the embryo depends on early in life while the seed germinates until the seedling has developed roots and sometimes leaves to start photosynthesis. 3.4.2 STRUCTURE Foliage Leaves Most leaves consist of a flat and thin structure called the leaf blade, or lamina, that is attached to the stem or branch by a narrow stalk called a petiole (Figure 3–12). The lamina may be simple or compound. Leaves lacking this stalk and attached directly to the stem are said to be sessile. The leaf is dorsoventral (flattened top to bottom), and as such the epidermis on the adaxial (upper) and abaxial (lower) surfaces experiences different environmental conditions. The upper epidermal layer of a leaf has a thicker cuticle than the lower layer. In some families, such as Portulacaceae and Begoniaceae, certain species have no stomata (openings or pores in the epidermis) on the adaxial epidermis. These pores are bordered by specialized cells called guard cells (Figure 3–13). In floating species of families such as Ranuculaceae, abaxial stomata are lacking; in completely submerged plants, such as certain species of Nymphaeaceae, stomata may be absent altogether. Most stomata occur on the abaxial surface, as do trichomes. The internal part of the leaf, the mesophyll, is equivalent to the cortex in the stem (Figure 3–14). Directly below the upper epidermis are columns of cells called palisade parenchyma. There may be FIGURE 3–12 A typical broad leaf. The size, shape, and other features vary widely among species. Lamina (leaf blade) Petiole Stipule FIGURE 3–13 (a) A surface view and (b) cross section of a stoma. Nucleus Stoma Guard cell Chloroplast Epidermal cell Guard cell Stoma Epidermis Mesophyll (a) Surface view (b) Cross section 76 Chapter 3 Plant Anatomy
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HORTICULTURE Principles and Practic
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HORTICULTURE Principles and Practic
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With love to Theresa, quarterback;
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Brief Contents Preface xxi PART 1 T
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Contents Preface xxi PART 1 THE UND
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5.3 PLANT GROWTH PROCESSES 160 5.4
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8.20 COMMON GREENHOUSE DISEASES 276
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12.3 INTERNAL ENVIRONMENTAL CONTROL
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PART 6 Summary 541 References and S
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22.18 INDOOR COMPOSTING SYSTEMS 668
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Preface Horticulture is the area of
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ACKNOWLEDGMENTS I am very grateful
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PART 1 THE UNDERLYING SCIENCE CHAPT
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1 What Is Horticulture? PURPOSE AND
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(a) (c) (b) (d) FIGURE 1-1 The many
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FIGURE 1 Bridge. The plaza view of
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CYCADS Many people mistake these pr
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FIGURE 2 The world's largest unbran
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FIGURE 2 Sold flowers are loaded on
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FIGURE 1-4 Formal landscaping featu
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1.4 ROLEOFTHENURSERY AND SEED INDUS
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1.5 HORTICULTURE AND SOCIETY Hortic
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TABLE 1-3 U.S. Horticultural Export
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Turfgrass Operation 1. Landscape te
Page 50 and 51: What Is Horticulture? This site pro
Page 52 and 53: Examples of botanical gardens http:
Page 54 and 55: 2 Classifying and Naming Horticultu
Page 56 and 57: Eight major taxa are commonly used
Page 58 and 59: TABLE 2-3 The Divisions of the King
Page 60 and 61: HISTORY OF PLANT TAXONOMY PAUL R. F
Page 62 and 63: AGE OF HERBALISTS Two major events
Page 64 and 65: possible system of nomenclature. Ho
Page 66 and 67: TABLE 1 Type Categories for Plant N
Page 68 and 69: 2.3 OTHER CLASSIFICATION SYSTEMS (O
Page 70 and 71: 2. Shrubs. A shrub has no main trun
Page 72 and 73: Simple Fruits Fleshy Fruits Drupe B
Page 74 and 75: FIGURE 2-14 A pome, represented by
Page 76 and 77: 2.3.5 CLASSIFICATION OF VEGETABLES
Page 78 and 79: (a) (b) FIGURE 2-22 (Source: George
Page 80 and 81: FIGURE 2-25 A narrowleaf plant. (So
Page 82 and 83: FIGURE 2-29 Parts of a typical gras
Page 84 and 85: such as rosemary, sage, thyme, marj
Page 86 and 87: c. Leaves d. Bulbs 2. Cut across (t
Page 88 and 89: Whole plant Organs FIGURE 3-1 Level
Page 90 and 91: ibonucleic acid (RNA), proteins, an
Page 92 and 93: called cristae; this extreme foldin
Page 94 and 95: By virtue of its position, the prim
Page 96 and 97: Phloem Tissue Structurally, phloem
Page 98 and 99: (a) Stalk (b) Culm FIGURE 3-5 Cross
Page 102 and 103: Upper epidermis Palisade layer FIGU
Page 104 and 105: usually occur in xerophytes. In cer
Page 106 and 107: FIGURE 3-22 Selected common leaf ma
Page 108 and 109: FIGURE 3-25 Selected common leaf ti
Page 110 and 111: absorption of water and minerals fr
Page 112 and 113: Outer bark Inner bark FIGURE 3-37 T
Page 114 and 115: Anther Filament Stamen FIGURE 3-41
Page 116 and 117: Exocarp Parts of a typi- FIGURE 3-4
Page 118 and 119: PRACTICAL EXPERIENCE LABORATORY 1.
Page 120 and 121: 4.1 CLIMATE, WEATHER, AND HORTICULT
Page 122 and 123: concentration in the atmosphere.A c
Page 124 and 125: TABLE 4-1 Climatic Adaptation of Se
Page 126 and 127: and upward. Another important gener
Page 128 and 129: Rate of photosynthesis mg/sq. dm/hr
Page 130 and 131: and plants that flower under only c
Page 132 and 133: times of the year. Growers start th
Page 134 and 135: content. This section is sometimes
Page 136 and 137: TABLE 4-7 Soil Mineral Nutrients Es
Page 138 and 139: Micronutrients (Trace Elements) Mic
Page 140 and 141: Neutral FIGURE 4-11 A representatio
Page 142 and 143: 4.4 FERTILIZERS Fertilizer sources
Page 144 and 145: One of the most commonly used contr
Page 146 and 147: Chlorosis (the yellowing of green l
Page 148 and 149: Fertilizers may be applied before p
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It is neither practical nor safe to
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Solution: How much of ammonium nitr
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1°C (34°F), the optimum temperatu
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Cellulose sponge Perched water tabl
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Overhead Sprinkler Irrigation Water
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FIGURE 4-19 Furrow irrigation of le
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can self-install an underground irr
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1. Surface drainage. Surface draina
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Secondary Tillage Primary tillage i
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(a) (b) (c) (d) FIGURE 4-20 (Source
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texture. The most commonly used gra
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TABLE 4-11 Selected Standard Mixes
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Steam Pasteurization Steam pasteuri
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Maracher, H. 1986. Mineral nutritio
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5 Plant Physiology PURPOSE AND EXPE
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Growth in an organism follows a cer
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5.1.2 THE ROLE OF SIGNALS IN GROWTH
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waxes are embedded. Waxes consist o
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5.3.1 PHOTOSYNTHESIS Photosynthesis
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CO 2 FIGURE 5-6 The C 4 pathway of
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Growth and Development The general
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TABLE 5-2 Energy Produced from Aero
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Certain plants are adapted to dry e
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conditions exist to sustain growth
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Shoot Elongation In certain plants,
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for success, since high temperature
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FIGURE 5-13 Ripening of plantain sh
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Terminal bud removed Unbranched pla
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conditions—pertaining to light, m
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them to maturity. The major process
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6 Breeding Horticultural Plants PUR
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Similarly, there can be no plant br
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APPLICATION, CHALLENGES, AND PROSPE
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hit with target DNA. Therefore, it
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Generally, within ten days of exper
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Aziz A.N., Sauve R.J., Zhou S., 200
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(b) F 1 Rr Rr round round F 2 RR R
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e.g., Aa × Aa), the lethal allele
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eeder’s equation. Simply stated,
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Before the seed or product becomes
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6.18.2 THE GENERAL STEPS OF RDNA TE
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2. Political disagreement. There ar
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REFERENCES AND SUGGESTED READING Ac
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PART 2 PROTECTING HORTICULTURAL PLA
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7 Biological Enemies of Horticultur
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8. Weeds may clog drains, waterways
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is also a root parasite that obtain
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LEGISLATIVE Both state and federal
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Example Integrated cultural, physic
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7.4.2 IMPORTANT INSECT ORDERS Insec
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Egg FIGURE 7-3 Life cycle of an ins
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sucking insects (also found with so
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FIGURE 7-12 Corn earworm damage. (S
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TABLE 7-1 Selected Fungal Diseases
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7.6.1 SMALL ANIMALS Rabbits, mice,
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FIGURE 7-16 The disease triangle. P
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fungitoxic exudates in its leaves,
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SUMMARY Insects are a major class o
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For the home growers or those who c
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for consumers and the environment).
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TABLE 8-1 Strategy 4: Strategies an
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gibberellic acid spray overcomes st
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In a competitive industry, a variet
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Chemicals gain access to humans thr
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2. Pesticide management. Controllin
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Every organism has its natural enem
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TABLE 8-3 Selected Examples of Biol
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1 2 YEAR 3 4 FIGURE 8-5 cycle. A cr
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6. Heat treatment. In the greenhous
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Organic Compounds (Organics) Organi
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under enclosed conditions (e.g., wa
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FIGURE 8-9 A tractor-mounted spraye
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8.11.9 LANDSCAPE PESTS AND THEIR CO
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application, a particular herbicide
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Further, they do not provide unifor
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SUMMARY Herbicides are chemicals us
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Sulfur may be applied for both prev
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8.23 PREVENTING GREENHOUSE DISEASES
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PART 3 PROPAGATING HORTICULTURAL PL
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9 Sexual Propagation PURPOSE AND EX
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Anther Microspore Megaspore mother
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Lettuce seeds Red light Darkness Fa
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FEDERAL AND STATE SEED LAWS Federal
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Germination Test In laboratory prac
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FIGURE 15 The essential structures
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processing into flour or meal). How
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physiologically immature seeds must
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seeds may be treated in this way be
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The two basic modes of seedling eme
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locations in the field. Home garden
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FIGURE 9-9 A plastic flat. (Source:
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(a) (b) FIGURE 9-12 (a) Sowing seed
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y the gardener or grower. Whatever
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REFERENCES AND SUGGESTED READING Co
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species enables vegetative propagat
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for rapid rooting. There are two ba
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Cutting involving one node (e.g., s
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This practice is especially importa
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10.6.4 STICKING THE CUTTING Cutting
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(a) Indexing by budding Diseased pl
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10.11 M ETHODS OF GRAFTING Grafting
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Scion Wax FIGURE 10-17 Steps in bar
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MODULE 3 BUDDING 10.12 TYPES OF BUD
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MODULE 4 LAYERING 10.13 TYPES OF LA
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Buried part of shoot is nicked FIGU
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variety of ways. In air layering, a
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FIGURE 10-34 by using cormels. Prop
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Psuedobulbs In the Dendrobium orchi
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The technique is used widely in cro
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PART 4 GROWING PLANTS INDOORS CHAPT
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11 Growing Houseplants PURPOSE AND
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TABLE 11-1 Common houseplants Commo
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Saddle leaf Philodendron selloum To
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Window Displays Plants in windows e
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CONTAINER GARDENS DR. TERRI W. STAR
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annuals and hardy perennial species
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of the large container filled with
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perfection about four to six weeks
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FIGURE 11-6 Flowers displayed on th
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TABLE 11-5 Plant Selected Plants fo
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The lighting condition near these w
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Fluorescent Lights Fluorescent ligh
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may be used for one pot or a group
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garden rooms, atriums, or a large c
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The photoperiod affects when the ho
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patted firm to keep the plant erect
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Other Materials Apart from clay and
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(a) ( FIGURE 11-25 Support for plan
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TABLE 11-7 Common Problems of House
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• Keep soil moist all the time
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• Prefers high temperatures • P
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amount and quality of light. If sup
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12 Controlled-Environment Horticult
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6. Curvilinear 7. Curved eave 8. Do
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Detached greenhouses have several a
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12.2.3 FRAME DESIGN There are two b
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horticultural business a less-expen
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Texas, Hawaii, and California. The
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source of heat for times when the t
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FIGURE 12-17 Greenhouse production
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FIGURE 12-21 Moving tables allowing
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Research program on greenhouse engi
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greenhouses equipped with a variety
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FIGURE 1 Annual energy required per
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This system was demonstrated in a 5
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FIGURE 6 Amounts of waste energy ut
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Ekholt, B.A., D.R. Mears, M.S. Gini
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or object to be warmed. Failure to
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objects in its path (e.g., the floo
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FIGURE 12-27 Motorized ventilation
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FIGURE 12-30 Movable internal shade
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FIGURE 12-33 A high pressure sodium
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Source of Water The quality of loca
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FIGURE 12-37 Overhead sprinkler irr
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Intermittent Feed Greenhouse plants
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However, in winter, greenhouse vent
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OUTCOMES ASSESSMENT 1. Explain the
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. Foliage or green plants. Foliage
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2. Labor. The size of the labor for
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FIGURE 13-1 Greenhouse production o
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FIGURE 13-2 Lettuce plug is inserte
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13.8.4 AGGREGATE HYDROPONIC SYSTEMS
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(a) (b) (c) FIGURE 13-6 Plug produc
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14 Growing Succulents PURPOSE AND E
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FIGURE 14-3 Leaf succulent represen
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frost-hardy. Their rosettes are usu
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TABLE 14-1 Plant Selected Popular S
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(a) (b) FIGURE 14-12 Typical bromel
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14.7.1 WHAT ARE CACTI? 14.7 CACTI C
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FIGURE 14-16 Opuntia. (Source: Crai
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FIGURE 14-23 Mammillaria. (Source:
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FIGURE 14-28 Both desert and jungle
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Growing mix Gravel Cacti (a) (b) FI
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PART5 GROWING PLANTS OUTDOORS: ORNA
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15 Principles of Landscaping PURPOS
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8. Create recreational grounds. Suc
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knowledge, with concern for resourc
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(a) (b) (c) FIGURE 15-2 The occurre
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GUIDELINES FOR LANDSCAPE DESIGN DAV
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the landscape. Some very successful
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Rhythm and Line Panoramic view of a
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FIGURE 15-10 A formal garden. The e
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How frequently do they entertain? A
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the patio should be located on the
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15.7.1 SELECTING PLANTS A homeowner
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Plant Arrangement in the Landscape
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Shadow FIGURE 15-15 Planting a tree
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SUMMARY Landscaping enhances the su
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3. Supply materials on a timely bas
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such as preparation rooms (for mixi
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of environmental fluctuations. Furt
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FIGURE 16-4 A bare-root tree seedli
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17 Installation of the Landscape PU
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for walks, driveways, and patios (F
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Planting may be limited to accentin
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17.3.3 PREPARING THE BED The soil s
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FIGURE 17-4 Bedding plants raised i
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SUMMARY Bedding plants are largely
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TABLE 17-6 Selected Ground Covers T
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TABLE 17-7 Selected Ornamental Gras
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they determine the success and surv
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12. Wildlife attraction. Trees in t
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pennsylvanica), hackberry (Celtis s
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y winds. A stake, which is often a
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TABLE 17-8 Selected Narrowleaf Ever
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TABLE 17-11 Selected Deciduous Shru
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Blooming bushes 1. Blue mist shrub
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Planting Bulblets and Bulbils Speci
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may be divided such that each secti
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FIGURE 18-1 (Source: George Acquaah
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Cool-Season (Temperate) Grasses In
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Growth Habit Turfgrasses are the mo
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Heavy Use Lawns on playgrounds and
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The seed should be free from weeds
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Source of Sod As with seed, sod sup
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A plug of sod FIGURE 18-7 Plugging
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way, plants are able to adapt to th
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form of a can placed on the lawn wi
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TABLE 18-3 Some Common Lawn and Tur
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even surface soil surface for layin
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MacCaskey, M. 1987. All about lawns
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Pruning is sometimes done in conjun
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4. Pruning may be done to reduce th
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19.4.2 SAWS A saw may be designed t
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defeat the purpose of pruning. The
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Bud withers as cut end dries back d
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19.6 STRATEGIES FOR PRUNING ABOVEGR
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Rejuvenation Pruning Cut canes to a
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3. In the third and subsequent year
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(a) Cut Prune (b) FIGURE 19-16 Step
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Eucalyptus and Paulownia are amenab
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TRAINING & PRUNING DECIDUOUS FRUIT
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Summer pruning eliminates an energy
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a) b) FIGURE 2 Newly planted apple
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FIGURE 6 Wooden limb spreaders can
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FIGURE 9. An apple tree trained to
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years to promote continued lateral
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Horizontal Espalier The horizontal
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19.16.1 CANE FRUITS Cane fruits are
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FIGURE 19-26 Shearing of Christmas
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pyramid-like form that is wider at
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After selecting the appropriate spe
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PART 6 GROWING PLANTS OUTDOORS: VEG
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20 Growing Vegetables Outdoors PURP
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The National Agricultural Statistic
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(This item omitted from WebBook edi
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growers should take to determine an
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pests and reduce/ eliminate hail da
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square yard (10 to 68 grams per squ
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High tunnels help increase the prof
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(This item omitted from WebBook edi
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20.4 VEGETABLE MARKET TYPES Fresh V
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Establishing the Crop Planting into
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home water supply from the tap. Thi
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Cole crop Cabbage Root Potato Bean
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6. Adequate nutrition. While overfe
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variable, ranging from creamy yello
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There are two general production pr
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This toxin is heat resistant and no
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large, or jumbo. The bulb may be sw
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REFERENCES Growing selected vegetab
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TABLE 21-1 Popular Herbs and Their
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(a) (b) (c) (d) (e) (f) FIGURE 21-1
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22 Organic Farming PURPOSE AND EXPE
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22.3 PRINCIPLES OF ORGANIC FARMING
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and the specific materials to be us
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22.8 MANAGING SOIL PHYSICAL QUALITY
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preemergent or early postemergent o
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Composting is a deliberate activity
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22.14.5 THE CARBON-TO-NITROGEN RATI
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Moisture Supply Water is required b
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Compost materials FIGURE 22-4 a wir
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As microbial decomposition proceeds
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Establishment and Management of an
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night, freezing can occur in spring
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accomplished by stratification. It
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transmitted by the dagger nematode
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PART 7 SPECIAL TECHNIQUES AND HANDL
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24 Cut Flowers and Floral Design PU
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to more than four-fold in standard
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Temperature and Humidity Wilting re
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FLORAL DESIGN: AN OVERVIEW BY WM. J
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Principle Definition Types (or Uses
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pH value-a measure of the acidity o
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FIGURE 6 Parallel Design-Parallel d
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24.3.2 TOOLS AND MATERIALS The tool
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3. Establish the focal point. 4. Ad
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Natural Drying To dry naturally, fl
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24.4.3 DRIED FLOWER ARRANGEMENTS Dr
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25 Terrarium Culture PURPOSE AND EX
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FIGURE 25-3 Terrarium containers ar
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FIGURE 25-5 Assortment of tools use
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25.6.7 ENHANCING THE DISPLAY Certai
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(a) (b) FIGURE 26-1 Bonsai can be c
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TABLE 26-3 Plant A Selection of Pop
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26.3.1 COLLECTING BONSAI PLANTS FRO
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Strip bark Bare branch FIGURE 26-9
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26.5.2 SANITATION It is critical to
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27 Postharvest Handling and Marketi
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whereas oranges are picked (they ha
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(b) (a) (c) (d) (e1) (e2) (f) FIGUR
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To reduce packaging injury, contain
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is replaced by the by-product of re
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Stored produce may lose some color,
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with pricing. When selling by volum
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(a) (b) FIGURE 27-5 Horticultural p
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APPENDIX A Temperature: Converting
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APPENDIX B Metric Conversion Chart
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APPENDIX D Common and Scientific Na
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Pecan (Carya illinoensis) Peony (Pa
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GLOSSARY A Abaxial Turned away from
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Cellulose A complex carbohydrate th
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Floriculture The science and practi
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M Macronutrient An essential elemen
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Root cap A mass of hard cells cover
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INDEX A-frame, 395 A-horizon, 108 A
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defined, 390 fertilization, 432-434
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Radiant heaters, 378 Radicle, 90 Re
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color plate 1 (a) (b) (c) (d) (e) M
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color plate 3 (b) (a) (c) (d) (e) (
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color plate 5 (a) (b) (d) (c) (e) (
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color plate 7 (b) (c) (d) (a) (e) (
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color plate 9 (a) (b) (c) (d) (e) (
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color plate 11 (a) (c) (b) (d) Grow
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color plate 13 (a) (b) (c) (d) (e)
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color plate 21 (a) (b) (c) (e) (d)
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color plate 23 (c) (b) (a) (d) (e)
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color plate 25 (c) (a) (b) (d) (e)
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color plate 27 (a1) (a2) (b2) (b1)
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color plate 31 (a) (b) (c) Floral d
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Horticulture Principles and Practices

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