Horticulture Principles and Practices

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Anther Microspore Megaspore mother cell FIGURE 9–1 Reproduction in flowering plants involves pollination and fertilization. Pollen grain Mature ovary Pollination Sperms Primary endosperm nucleus (3n) Zygote (2n) Fertilization into the endosperm, a nutrient-rich mass of cells that provides nourishment to the embryo until it develops to a stage where it becomes self-supporting (seedling stage). The mature ovule is the seed. The outer tissue of the ovules (the inner and outer integuments) fuse and lose their water to become the seed coat, or testa. The endosperm contains large amounts of starch that serve as food reserves to be broken down into glucose for use during seed germination. In monocots, the outermost layer of the endosperm is called the aleurone layer and is the site of protein storage. In seeds such as sunflower, mustard, pine, and fir, the endosperm stores a large amount of fats and oils. The embryo is variable in size in terms of the volume of the seed it occupies. It occupies a large volume in species such as pea and oak, but occupies a relatively small volume in cereals. In species such as holly and orchid, the embryo remains relatively undifferentiated. However, in grass, the embryo is highly differentiated into structures including the shoot apex, scutellum (single cotyledon), and coleoptile (a cylindrical protective leaf). The root apex is protected by the coleorrhiza. Seed The mature ovule of a flowering plant. 9.2 SEED PRODUCTION AND CERTIFICATION PROCESS Plant breeders are engaged in plant improvement. Old cultivars are changed by importing new genes that condition improved characteristics, making new cultivars more disease resistant, higher yielding, more beautiful, and better in other ways according to breeding objectives. Once the breeder has completed the breeding program and tested the new material extensively, the seed is then released by the researchers to special producers for propagation. Before the seed becomes available to the ordinary grower, it goes through stages of increase and certification. According to the stages, there are four classes of seed (Figure 9–2): 1. Breeder seed. A small amount of seed is developed and released by a breeder as the source of foundation seed. 2. Foundation seed. Breeder seed is increased under supervision of agricultural research stations and monitored for genetic purity and identity. 9.2 Seed Production and Certification Process 285
FIGURE 9–2 Commercial classes of seed. The differences depend on the quantities of seed handled and the person responsible for seed maintenance. BREEDER SEED Small quantity of original seed produced by plant breeders through a carefully designed and conducted breeding program and according to specific objectives. Controlled by breeder/ institution or company. FOUNDATION SEED An increase of the breeder seed by Agricultural Experimental Stations or reputable growers under supervision of researchers. REGISTERED SEED An increase of foundation seed by registered growers. CERTIFIED SEED An increase of registered seed (or foundation seed) that is inspected and approved by certifying agency for sale to farmers and other growers. Certified Seed The progeny of registered seed that is maintained at a satisfactory genetic identity and purity, and approved and certified by an official certifying agency. 3. Registered seed. Foundation seed is distributed to certified seed growers to be further increased for distribution. 4. Certified seed. The progeny of registered seed is sold to farmers. During the process of increase, certifying agencies in the state or region of production monitor the activity to ensure that the product meets standards set for the crop. Before sale, the certified seed grower is required to perform certain basic analyses (seed quality analyses) of the seed as described in the following section. 9.3 SEED LAW The seed law is an act intended to regulate the labeling, coloring, sale, offering, exposing, or transporting for sale of agricultural, vegetable, flower, and tree and shrub seeds, to prevent those in the industry from misrepresenting their products to customers. It is important to assure that seed offered for sale to growers of agricultural crops, landscapers, and private homeowners meets quality standards set by the state in which the products are used or produced. The Association of American Seed Control Officials (AASCO), comprising officials from the United States and Canada, developed the Recommended Uniform State Seed Law (RUSSL) as a model law for states and federal programs. The RUSSL is to be enforced only after a state has enacted it into its own seed law by its legislature. The organization meets annually to be updated on new developments in the seed industry and to update the RUSSL as needed. Among other things, a seed law describes the issue of seed tags and what specific information goes on them, brand registration, seed permits, arbitration of seed complaints, and the enforcement of the law. 286 Chapter 9 Sexual Propagation 9.4 SEED QUALITY ANALYSIS Certified seed producers are required to declare their seed analysis on the seed packet or tag. The interpretation of the tag is provided in Table 9–1. To do so, a seed test must be conducted. The common seed tests available in the seed industry include the germination test, cold test, tetrazolium test, and purity test.
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HORTICULTURE Principles and Practic
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HORTICULTURE Principles and Practic
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With love to Theresa, quarterback;
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Brief Contents Preface xxi PART 1 T
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Contents Preface xxi PART 1 THE UND
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5.3 PLANT GROWTH PROCESSES 160 5.4
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8.20 COMMON GREENHOUSE DISEASES 276
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12.3 INTERNAL ENVIRONMENTAL CONTROL
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PART 6 Summary 541 References and S
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22.18 INDOOR COMPOSTING SYSTEMS 668
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Preface Horticulture is the area of
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ACKNOWLEDGMENTS I am very grateful
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PART 1 THE UNDERLYING SCIENCE CHAPT
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1 What Is Horticulture? PURPOSE AND
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(a) (c) (b) (d) FIGURE 1-1 The many
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FIGURE 1 Bridge. The plaza view of
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CYCADS Many people mistake these pr
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FIGURE 2 The world's largest unbran
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FIGURE 2 Sold flowers are loaded on
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FIGURE 1-4 Formal landscaping featu
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1.4 ROLEOFTHENURSERY AND SEED INDUS
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1.5 HORTICULTURE AND SOCIETY Hortic
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TABLE 1-3 U.S. Horticultural Export
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Turfgrass Operation 1. Landscape te
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What Is Horticulture? This site pro
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Examples of botanical gardens http:
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2 Classifying and Naming Horticultu
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Eight major taxa are commonly used
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TABLE 2-3 The Divisions of the King
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HISTORY OF PLANT TAXONOMY PAUL R. F
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AGE OF HERBALISTS Two major events
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possible system of nomenclature. Ho
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TABLE 1 Type Categories for Plant N
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2.3 OTHER CLASSIFICATION SYSTEMS (O
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2. Shrubs. A shrub has no main trun
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Simple Fruits Fleshy Fruits Drupe B
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FIGURE 2-14 A pome, represented by
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2.3.5 CLASSIFICATION OF VEGETABLES
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(a) (b) FIGURE 2-22 (Source: George
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FIGURE 2-25 A narrowleaf plant. (So
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FIGURE 2-29 Parts of a typical gras
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such as rosemary, sage, thyme, marj
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c. Leaves d. Bulbs 2. Cut across (t
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Whole plant Organs FIGURE 3-1 Level
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ibonucleic acid (RNA), proteins, an
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called cristae; this extreme foldin
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By virtue of its position, the prim
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Phloem Tissue Structurally, phloem
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(a) Stalk (b) Culm FIGURE 3-5 Cross
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Scale Compressed stem (a) Whole bul
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Upper epidermis Palisade layer FIGU
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usually occur in xerophytes. In cer
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FIGURE 3-22 Selected common leaf ma
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FIGURE 3-25 Selected common leaf ti
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absorption of water and minerals fr
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Outer bark Inner bark FIGURE 3-37 T
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Anther Filament Stamen FIGURE 3-41
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Exocarp Parts of a typi- FIGURE 3-4
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PRACTICAL EXPERIENCE LABORATORY 1.
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4.1 CLIMATE, WEATHER, AND HORTICULT
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concentration in the atmosphere.A c
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TABLE 4-1 Climatic Adaptation of Se
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and upward. Another important gener
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Rate of photosynthesis mg/sq. dm/hr
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and plants that flower under only c
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times of the year. Growers start th
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content. This section is sometimes
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TABLE 4-7 Soil Mineral Nutrients Es
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Micronutrients (Trace Elements) Mic
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Neutral FIGURE 4-11 A representatio
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4.4 FERTILIZERS Fertilizer sources
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One of the most commonly used contr
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Chlorosis (the yellowing of green l
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Fertilizers may be applied before p
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It is neither practical nor safe to
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Solution: How much of ammonium nitr
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1°C (34°F), the optimum temperatu
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Cellulose sponge Perched water tabl
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Overhead Sprinkler Irrigation Water
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FIGURE 4-19 Furrow irrigation of le
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can self-install an underground irr
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1. Surface drainage. Surface draina
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Secondary Tillage Primary tillage i
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(a) (b) (c) (d) FIGURE 4-20 (Source
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texture. The most commonly used gra
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TABLE 4-11 Selected Standard Mixes
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Steam Pasteurization Steam pasteuri
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Maracher, H. 1986. Mineral nutritio
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5 Plant Physiology PURPOSE AND EXPE
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Growth in an organism follows a cer
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5.1.2 THE ROLE OF SIGNALS IN GROWTH
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waxes are embedded. Waxes consist o
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5.3.1 PHOTOSYNTHESIS Photosynthesis
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CO 2 FIGURE 5-6 The C 4 pathway of
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Growth and Development The general
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TABLE 5-2 Energy Produced from Aero
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Certain plants are adapted to dry e
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conditions exist to sustain growth
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Shoot Elongation In certain plants,
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for success, since high temperature
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FIGURE 5-13 Ripening of plantain sh
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Terminal bud removed Unbranched pla
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conditions—pertaining to light, m
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them to maturity. The major process
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6 Breeding Horticultural Plants PUR
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Similarly, there can be no plant br
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APPLICATION, CHALLENGES, AND PROSPE
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hit with target DNA. Therefore, it
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Generally, within ten days of exper
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Aziz A.N., Sauve R.J., Zhou S., 200
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(b) F 1 Rr Rr round round F 2 RR R
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e.g., Aa × Aa), the lethal allele
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eeder’s equation. Simply stated,
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Before the seed or product becomes
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6.18.2 THE GENERAL STEPS OF RDNA TE
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2. Political disagreement. There ar
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REFERENCES AND SUGGESTED READING Ac
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PART 2 PROTECTING HORTICULTURAL PLA
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7 Biological Enemies of Horticultur
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8. Weeds may clog drains, waterways
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is also a root parasite that obtain
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LEGISLATIVE Both state and federal
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Example Integrated cultural, physic
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7.4.2 IMPORTANT INSECT ORDERS Insec
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Egg FIGURE 7-3 Life cycle of an ins
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sucking insects (also found with so
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FIGURE 7-12 Corn earworm damage. (S
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TABLE 7-1 Selected Fungal Diseases
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7.6.1 SMALL ANIMALS Rabbits, mice,
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Page 264 and 265: SUMMARY Insects are a major class o
Page 266 and 267: For the home growers or those who c
Page 268 and 269: for consumers and the environment).
Page 270 and 271: TABLE 8-1 Strategy 4: Strategies an
Page 272 and 273: gibberellic acid spray overcomes st
Page 274 and 275: In a competitive industry, a variet
Page 276 and 277: Chemicals gain access to humans thr
Page 278 and 279: 2. Pesticide management. Controllin
Page 280 and 281: Every organism has its natural enem
Page 282 and 283: TABLE 8-3 Selected Examples of Biol
Page 284 and 285: 1 2 YEAR 3 4 FIGURE 8-5 cycle. A cr
Page 286 and 287: 6. Heat treatment. In the greenhous
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Page 290 and 291: under enclosed conditions (e.g., wa
Page 292 and 293: FIGURE 8-9 A tractor-mounted spraye
Page 294 and 295: 8.11.9 LANDSCAPE PESTS AND THEIR CO
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Page 298 and 299: Further, they do not provide unifor
Page 300 and 301: SUMMARY Herbicides are chemicals us
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Page 306 and 307: PART 3 PROPAGATING HORTICULTURAL PL
Page 308 and 309: 9 Sexual Propagation PURPOSE AND EX
Page 312 and 313: Lettuce seeds Red light Darkness Fa
Page 314 and 315: FEDERAL AND STATE SEED LAWS Federal
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Page 318 and 319: FIGURE 15 The essential structures
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Page 324 and 325: seeds may be treated in this way be
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Page 330 and 331: FIGURE 9-9 A plastic flat. (Source:
Page 332 and 333: (a) (b) FIGURE 9-12 (a) Sowing seed
Page 334 and 335: y the gardener or grower. Whatever
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Page 338 and 339: species enables vegetative propagat
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Page 342 and 343: Cutting involving one node (e.g., s
Page 344 and 345: This practice is especially importa
Page 346 and 347: 10.6.4 STICKING THE CUTTING Cutting
Page 348 and 349: (a) Indexing by budding Diseased pl
Page 350 and 351: 10.11 M ETHODS OF GRAFTING Grafting
Page 352 and 353: Scion Wax FIGURE 10-17 Steps in bar
Page 354 and 355: MODULE 3 BUDDING 10.12 TYPES OF BUD
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variety of ways. In air layering, a
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FIGURE 10-34 by using cormels. Prop
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Psuedobulbs In the Dendrobium orchi
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The technique is used widely in cro
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PART 4 GROWING PLANTS INDOORS CHAPT
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11 Growing Houseplants PURPOSE AND
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TABLE 11-1 Common houseplants Commo
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Saddle leaf Philodendron selloum To
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Window Displays Plants in windows e
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CONTAINER GARDENS DR. TERRI W. STAR
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annuals and hardy perennial species
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of the large container filled with
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perfection about four to six weeks
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FIGURE 11-6 Flowers displayed on th
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TABLE 11-5 Plant Selected Plants fo
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The lighting condition near these w
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Fluorescent Lights Fluorescent ligh
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may be used for one pot or a group
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garden rooms, atriums, or a large c
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The photoperiod affects when the ho
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patted firm to keep the plant erect
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Other Materials Apart from clay and
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(a) ( FIGURE 11-25 Support for plan
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TABLE 11-7 Common Problems of House
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• Keep soil moist all the time
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• Prefers high temperatures • P
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amount and quality of light. If sup
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12 Controlled-Environment Horticult
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6. Curvilinear 7. Curved eave 8. Do
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Detached greenhouses have several a
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12.2.3 FRAME DESIGN There are two b
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horticultural business a less-expen
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Texas, Hawaii, and California. The
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source of heat for times when the t
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FIGURE 12-17 Greenhouse production
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FIGURE 12-21 Moving tables allowing
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Research program on greenhouse engi
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greenhouses equipped with a variety
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FIGURE 1 Annual energy required per
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This system was demonstrated in a 5
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FIGURE 6 Amounts of waste energy ut
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Ekholt, B.A., D.R. Mears, M.S. Gini
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or object to be warmed. Failure to
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objects in its path (e.g., the floo
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FIGURE 12-27 Motorized ventilation
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FIGURE 12-30 Movable internal shade
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FIGURE 12-33 A high pressure sodium
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Source of Water The quality of loca
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FIGURE 12-37 Overhead sprinkler irr
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Intermittent Feed Greenhouse plants
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However, in winter, greenhouse vent
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OUTCOMES ASSESSMENT 1. Explain the
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. Foliage or green plants. Foliage
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2. Labor. The size of the labor for
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FIGURE 13-1 Greenhouse production o
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FIGURE 13-2 Lettuce plug is inserte
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13.8.4 AGGREGATE HYDROPONIC SYSTEMS
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(a) (b) (c) FIGURE 13-6 Plug produc
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14 Growing Succulents PURPOSE AND E
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FIGURE 14-3 Leaf succulent represen
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frost-hardy. Their rosettes are usu
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TABLE 14-1 Plant Selected Popular S
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(a) (b) FIGURE 14-12 Typical bromel
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14.7.1 WHAT ARE CACTI? 14.7 CACTI C
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FIGURE 14-16 Opuntia. (Source: Crai
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FIGURE 14-23 Mammillaria. (Source:
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FIGURE 14-28 Both desert and jungle
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Growing mix Gravel Cacti (a) (b) FI
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PART5 GROWING PLANTS OUTDOORS: ORNA
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15 Principles of Landscaping PURPOS
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8. Create recreational grounds. Suc
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knowledge, with concern for resourc
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(a) (b) (c) FIGURE 15-2 The occurre
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GUIDELINES FOR LANDSCAPE DESIGN DAV
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the landscape. Some very successful
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Rhythm and Line Panoramic view of a
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FIGURE 15-10 A formal garden. The e
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How frequently do they entertain? A
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the patio should be located on the
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15.7.1 SELECTING PLANTS A homeowner
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Plant Arrangement in the Landscape
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Shadow FIGURE 15-15 Planting a tree
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SUMMARY Landscaping enhances the su
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3. Supply materials on a timely bas
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such as preparation rooms (for mixi
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of environmental fluctuations. Furt
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FIGURE 16-4 A bare-root tree seedli
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17 Installation of the Landscape PU
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for walks, driveways, and patios (F
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Planting may be limited to accentin
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17.3.3 PREPARING THE BED The soil s
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FIGURE 17-4 Bedding plants raised i
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SUMMARY Bedding plants are largely
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TABLE 17-6 Selected Ground Covers T
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TABLE 17-7 Selected Ornamental Gras
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they determine the success and surv
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12. Wildlife attraction. Trees in t
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pennsylvanica), hackberry (Celtis s
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y winds. A stake, which is often a
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TABLE 17-8 Selected Narrowleaf Ever
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TABLE 17-11 Selected Deciduous Shru
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Blooming bushes 1. Blue mist shrub
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Planting Bulblets and Bulbils Speci
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may be divided such that each secti
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FIGURE 18-1 (Source: George Acquaah
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Cool-Season (Temperate) Grasses In
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Growth Habit Turfgrasses are the mo
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Heavy Use Lawns on playgrounds and
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The seed should be free from weeds
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Source of Sod As with seed, sod sup
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A plug of sod FIGURE 18-7 Plugging
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way, plants are able to adapt to th
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form of a can placed on the lawn wi
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TABLE 18-3 Some Common Lawn and Tur
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even surface soil surface for layin
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MacCaskey, M. 1987. All about lawns
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Pruning is sometimes done in conjun
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4. Pruning may be done to reduce th
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19.4.2 SAWS A saw may be designed t
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defeat the purpose of pruning. The
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Bud withers as cut end dries back d
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19.6 STRATEGIES FOR PRUNING ABOVEGR
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Rejuvenation Pruning Cut canes to a
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3. In the third and subsequent year
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(a) Cut Prune (b) FIGURE 19-16 Step
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Eucalyptus and Paulownia are amenab
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TRAINING & PRUNING DECIDUOUS FRUIT
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Summer pruning eliminates an energy
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a) b) FIGURE 2 Newly planted apple
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FIGURE 6 Wooden limb spreaders can
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FIGURE 9. An apple tree trained to
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years to promote continued lateral
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Horizontal Espalier The horizontal
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19.16.1 CANE FRUITS Cane fruits are
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FIGURE 19-26 Shearing of Christmas
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pyramid-like form that is wider at
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After selecting the appropriate spe
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PART 6 GROWING PLANTS OUTDOORS: VEG
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20 Growing Vegetables Outdoors PURP
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The National Agricultural Statistic
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(This item omitted from WebBook edi
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growers should take to determine an
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pests and reduce/ eliminate hail da
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square yard (10 to 68 grams per squ
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High tunnels help increase the prof
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(This item omitted from WebBook edi
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20.4 VEGETABLE MARKET TYPES Fresh V
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Establishing the Crop Planting into
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home water supply from the tap. Thi
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Cole crop Cabbage Root Potato Bean
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6. Adequate nutrition. While overfe
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variable, ranging from creamy yello
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There are two general production pr
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This toxin is heat resistant and no
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large, or jumbo. The bulb may be sw
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REFERENCES Growing selected vegetab
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TABLE 21-1 Popular Herbs and Their
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(a) (b) (c) (d) (e) (f) FIGURE 21-1
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22 Organic Farming PURPOSE AND EXPE
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22.3 PRINCIPLES OF ORGANIC FARMING
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and the specific materials to be us
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22.8 MANAGING SOIL PHYSICAL QUALITY
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preemergent or early postemergent o
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Composting is a deliberate activity
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22.14.5 THE CARBON-TO-NITROGEN RATI
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Moisture Supply Water is required b
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Compost materials FIGURE 22-4 a wir
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As microbial decomposition proceeds
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Establishment and Management of an
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night, freezing can occur in spring
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accomplished by stratification. It
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transmitted by the dagger nematode
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PART 7 SPECIAL TECHNIQUES AND HANDL
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24 Cut Flowers and Floral Design PU
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to more than four-fold in standard
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Temperature and Humidity Wilting re
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FLORAL DESIGN: AN OVERVIEW BY WM. J
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Principle Definition Types (or Uses
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pH value-a measure of the acidity o
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FIGURE 6 Parallel Design-Parallel d
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24.3.2 TOOLS AND MATERIALS The tool
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3. Establish the focal point. 4. Ad
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Natural Drying To dry naturally, fl
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24.4.3 DRIED FLOWER ARRANGEMENTS Dr
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25 Terrarium Culture PURPOSE AND EX
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FIGURE 25-3 Terrarium containers ar
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FIGURE 25-5 Assortment of tools use
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25.6.7 ENHANCING THE DISPLAY Certai
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(a) (b) FIGURE 26-1 Bonsai can be c
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TABLE 26-3 Plant A Selection of Pop
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26.3.1 COLLECTING BONSAI PLANTS FRO
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Strip bark Bare branch FIGURE 26-9
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26.5.2 SANITATION It is critical to
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27 Postharvest Handling and Marketi
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whereas oranges are picked (they ha
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(b) (a) (c) (d) (e1) (e2) (f) FIGUR
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To reduce packaging injury, contain
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is replaced by the by-product of re
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Stored produce may lose some color,
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with pricing. When selling by volum
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(a) (b) FIGURE 27-5 Horticultural p
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APPENDIX A Temperature: Converting
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APPENDIX B Metric Conversion Chart
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APPENDIX D Common and Scientific Na
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Pecan (Carya illinoensis) Peony (Pa
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GLOSSARY A Abaxial Turned away from
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Cellulose A complex carbohydrate th
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Floriculture The science and practi
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M Macronutrient An essential elemen
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Root cap A mass of hard cells cover
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INDEX A-frame, 395 A-horizon, 108 A
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defined, 390 fertilization, 432-434
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Radiant heaters, 378 Radicle, 90 Re
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color plate 1 (a) (b) (c) (d) (e) M
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color plate 3 (b) (a) (c) (d) (e) (
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color plate 5 (a) (b) (d) (c) (e) (
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color plate 7 (b) (c) (d) (a) (e) (
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color plate 9 (a) (b) (c) (d) (e) (
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color plate 11 (a) (c) (b) (d) Grow
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color plate 13 (a) (b) (c) (d) (e)
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color plate 21 (a) (b) (c) (e) (d)
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color plate 23 (c) (b) (a) (d) (e)
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color plate 25 (c) (a) (b) (d) (e)
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color plate 27 (a1) (a2) (b2) (b1)
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color plate 31 (a) (b) (c) Floral d
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Horticulture Principles and Practices

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