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_9t<br />
Volcanic type islands built by volcenic activity and therefore with substrateg derived from<br />
lava (usuelly basalt) and volcanic ash.<br />
Ebvated reefg - ielande or parts of islandg compoeed of raised coral platforms or limestone.<br />
Lcnr islande - eomposed of sand and coral rubble accumulated m<br />
sea-level. This is the typical type m atolls and barrier<br />
occutr es coastal or beach areas on other island types.<br />
The islands in turn rnay be flanked by one or a eombination of several reef structures.<br />
Following Dahl (1980), these may be described as follows. An algal reef is a calcareoug<br />
structure in which corelline algae are principal eontributors to reef Lonstruction end s:rface<br />
eover' A coral reef, m the other hand, is a structure actively constructed by the skeletal<br />
deposition of hermatypic corals and associated organisms. Corai reefe in turn take the form<br />
of:<br />
Atoll reefs - annular reefs generally with an internal lagoon unassociated with any major<br />
landmass.<br />
Barrier reefs - reefs offshore from a major land mass and<br />
or navigable channel.<br />
separaled from it by a deep lagoon<br />
Fringing reefs - reefs growing directly out fnom the coastline and not separated from it by<br />
more than a shallow depression.<br />
Lagoon reefs or patch reefs - reef structures developing in the sheltered waters of a lagoon.<br />
A rrnon-growing" reef is a calcareous structure now covered with organisms not<br />
contributing significantly to skeletal accumulation or reef qrowth. Subrnerged reefs are those<br />
"drowned" by zubsidence to depths below which reef qro;tn has been insufficient to regain<br />
the zurface.<br />
.<br />
Similarly, the marine environment may be characterized by different structural criteria<br />
(Dahl, 1980). Substrates in different areas may be rocky (calcareous or non-calcareous) or<br />
uneonsolidated. In addition, reef community structure would depend on whether the island is<br />
subrnerging, emerging or +parently stationary relative to sea level, and whether the setting<br />
is exposed or protecLed with respect to lhe physical forces of lhe sea. These factors are all<br />
part of lhe variability of the physical and chemical environnent of island ecosystems (whicfr<br />
will be discussed in the following section), and which contribuLe signif icantly to the<br />
uniqueness of each island.<br />
,<br />
a reef platform at or near<br />
reefs, and also frequently<br />
Descriptions of reef structune and nnrphology in the tropical Pacific are relatively<br />
few. ssne studies that may be nrentioned are thosl m lhe cook lslands (Gibbs et al., r97l;<br />
Stoddart and Pillai, I97J)i Fanning lsland (Maragos, 1974a and b; Roy and Shith,-197I);<br />
French Polvnesia (chevalier, 1975b; Ricard 4 4.; thl); rhe Lau tslandi in Fiji (phipps and<br />
Preobrazhensky, 1975); the Marshall Islands-(L-acid, 1975; MacNeil, Lgiz)i the New Hebrides<br />
(now Vanuatu) in M_elanesia (Guilcher, 1974); New caledonia (chevalier, '197)a); p+ua r.rlw<br />
Guinea (Weber, I975b; Whitehouse, I97)); Samoa (Mayor, L9Z4); and 'the iotornon Islands<br />
(Jmes, 1977; Morton, 1974; Weber, L97Ja).<br />
The physical structure of an island is a major determinant of the types of ecosystems<br />
present (Dahl' f980)- A elear example may be found in the case of islands wilh exposed end<br />
protected sides, such as atolls. Coral reefs tend to be npre robust and better developed on<br />
the exposed sides due to rnore adequate wat.er circulation and greater availabiiity of<br />
nutrients. In addition, reef forms and lagoon types are determined by the structure and<br />
history of the island substrate (Dahl, I9S0). In general, Lherefore, a broad distinction can be<br />
made among continental islands, high volcanic islands, elevated reefs, and low coral islands,<br />
but it is also necessary to consider the delailed origin and structure of each, and its<br />
geographic location in terms of the origin and evolution of flora and fauna (E1io[t, Lg;-J).<br />
Dasmann (1971) attributes lhe uniqueness of island ecosystems to their relative isolation<br />
which induces peculiar evolutionary patlerns in each separate setting. Differences among<br />
similarly situated neef habitats are manifested, for example, in t6e differing rates oi<br />
productiviiy of at.oll lagoons in the South pacific (Ricard and belesalle, lggl).