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_9t<br />

Volcanic type islands built by volcenic activity and therefore with substrateg derived from<br />

lava (usuelly basalt) and volcanic ash.<br />

Ebvated reefg - ielande or parts of islandg compoeed of raised coral platforms or limestone.<br />

Lcnr islande - eomposed of sand and coral rubble accumulated m<br />

sea-level. This is the typical type m atolls and barrier<br />

occutr es coastal or beach areas on other island types.<br />

The islands in turn rnay be flanked by one or a eombination of several reef structures.<br />

Following Dahl (1980), these may be described as follows. An algal reef is a calcareoug<br />

structure in which corelline algae are principal eontributors to reef Lonstruction end s:rface<br />

eover' A coral reef, m the other hand, is a structure actively constructed by the skeletal<br />

deposition of hermatypic corals and associated organisms. Corai reefe in turn take the form<br />

of:<br />

Atoll reefs - annular reefs generally with an internal lagoon unassociated with any major<br />

landmass.<br />

Barrier reefs - reefs offshore from a major land mass and<br />

or navigable channel.<br />

separaled from it by a deep lagoon<br />

Fringing reefs - reefs growing directly out fnom the coastline and not separated from it by<br />

more than a shallow depression.<br />

Lagoon reefs or patch reefs - reef structures developing in the sheltered waters of a lagoon.<br />

A rrnon-growing" reef is a calcareous structure now covered with organisms not<br />

contributing significantly to skeletal accumulation or reef qrowth. Subrnerged reefs are those<br />

"drowned" by zubsidence to depths below which reef qro;tn has been insufficient to regain<br />

the zurface.<br />

.<br />

Similarly, the marine environment may be characterized by different structural criteria<br />

(Dahl, 1980). Substrates in different areas may be rocky (calcareous or non-calcareous) or<br />

uneonsolidated. In addition, reef community structure would depend on whether the island is<br />

subrnerging, emerging or +parently stationary relative to sea level, and whether the setting<br />

is exposed or protecLed with respect to lhe physical forces of lhe sea. These factors are all<br />

part of lhe variability of the physical and chemical environnent of island ecosystems (whicfr<br />

will be discussed in the following section), and which contribuLe signif icantly to the<br />

uniqueness of each island.<br />

,<br />

a reef platform at or near<br />

reefs, and also frequently<br />

Descriptions of reef structune and nnrphology in the tropical Pacific are relatively<br />

few. ssne studies that may be nrentioned are thosl m lhe cook lslands (Gibbs et al., r97l;<br />

Stoddart and Pillai, I97J)i Fanning lsland (Maragos, 1974a and b; Roy and Shith,-197I);<br />

French Polvnesia (chevalier, 1975b; Ricard 4 4.; thl); rhe Lau tslandi in Fiji (phipps and<br />

Preobrazhensky, 1975); the Marshall Islands-(L-acid, 1975; MacNeil, Lgiz)i the New Hebrides<br />

(now Vanuatu) in M_elanesia (Guilcher, 1974); New caledonia (chevalier, '197)a); p+ua r.rlw<br />

Guinea (Weber, I975b; Whitehouse, I97)); Samoa (Mayor, L9Z4); and 'the iotornon Islands<br />

(Jmes, 1977; Morton, 1974; Weber, L97Ja).<br />

The physical structure of an island is a major determinant of the types of ecosystems<br />

present (Dahl' f980)- A elear example may be found in the case of islands wilh exposed end<br />

protected sides, such as atolls. Coral reefs tend to be npre robust and better developed on<br />

the exposed sides due to rnore adequate wat.er circulation and greater availabiiity of<br />

nutrients. In addition, reef forms and lagoon types are determined by the structure and<br />

history of the island substrate (Dahl, I9S0). In general, Lherefore, a broad distinction can be<br />

made among continental islands, high volcanic islands, elevated reefs, and low coral islands,<br />

but it is also necessary to consider the delailed origin and structure of each, and its<br />

geographic location in terms of the origin and evolution of flora and fauna (E1io[t, Lg;-J).<br />

Dasmann (1971) attributes lhe uniqueness of island ecosystems to their relative isolation<br />

which induces peculiar evolutionary patlerns in each separate setting. Differences among<br />

similarly situated neef habitats are manifested, for example, in t6e differing rates oi<br />

productiviiy of at.oll lagoons in the South pacific (Ricard and belesalle, lggl).

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