9. Tribe ANTHEMIDEAE

752
5. Cancrinia tianschanica (Krascheninnikov) Tzvelev in
Schischkin & Bobrov, Fl. URSS 26: 315. 1961.
天山小甘菊 tian shan xiao gan ju
Cancrinia chrysocephala Karelin & Kirilov subsp. tianschanica
Krascheninnikov, Bot. Mater. Gerb. Glavn. Bot. Sada
R.S.F.S.R. 3: 81. 1922.
Herbs, perennial, ± scapose, to 10 cm tall, sparsely grayish
hirsute, hairs simple. Basal leaves many, petiolate; leaf blade
oblong or ovate, 2–3 × 0.5–1 cm, 2-pinnatipartite, both surfaces
densely lanate; primary lateral lobes 3- or 4-paired; ultimate
ANTHEMIDEAE
lobes obtuse or slightly acute at apex. Stem leaves very reduced,
margin entire. Capitulum solitary, long pedunculate. Involucres
coryliform, 4–6 mm in diam.; phyllaries in 3 rows,
herbaceous, scarious margin narrow, glossy brown, outer phyllaries
lanceolate or linear-lanceolate, inner ones longer, oblonglinear.
Florets yellow, 5-lobed. Achenes 1.5–2.2 mm, 5- or 6ribbed,
sparsely pilose. Corona scales 8–12, linear or ellipticlinear.
Fl. and fr. Jul–Aug.
Rocky places on mountain slopes, meadows, gravelly places,
floodlands; ca. 3200 m. Xinjiang (Tian Shan) [Kazakhstan (C Tian
Shan)].
158. RICHTERIA Karelin & Kirilov, Bull. Soc. Imp. Naturalistes Moscou 15: 126. 1842.
灰叶匹菊属 hui ye pi ju shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Subshrubs, basally woody; most parts tomentose; stems covered by sheathing leaf bases. Leaves alternate and basally rosulate,
blade pinnatisect. Capitula solitary, pedunculate, radiate. Phyllaries with dark brown margins. Receptacle convex, epaleate. Ray florets
female; lamina white. Disk corolla 5-lobed. Achenes faintly 6–10-ribbed, with sessile glands. Pappus of 6–10 obovate, apically
brownish scales at least 1/2 as long as corolla.
Three species: Afghanistan, NW China, N India, Mongolia, Russia; C Asia, SW Asia (Iran); one species in China.
1. Richteria pyrethroides Karelin & Kirilov, Bull. Soc. Imp.
Naturalistes Moscou 15: 127. 1842.
灰叶匹菊 hui ye pi ju
Chrysanthemum arassanicum C. Winkler; C. artemisiifolium
Klatt; C. pyrethroides (Karelin & Kirilov) B. Fedtschenko;
C. richteria Bentham; C. transiliense (Herder) Minkwitz;
Pyrethrum arassanicum (C. Winkler) O. Fedtschenko; P.
pyrethroides (Karelin & Kirilov) B. Fedtschenko ex Krascheninnikov;
P. transiliense (Herder) Regel; P. transiliense var. subsessile
Regel; P. transiliense var. subvillosum Regel; P. transiliense
var. tomentosum Regel; Tanacetum transiliense Herder.
Herbs, perennial, (5–)10–40 cm tall, rhizomatous; stems
erect or ascending, base covered with sheathing leaf bases, fascicled,
rarely solitary, few branched or not, gray-white, villous
with curved long simple hairs especially below capitulum. Bas-
al leaves: petioles to 4 cm; leaf blade narrowly elliptic, 1.5–7 ×
0.6–2 cm, 2-pinnatisect, both surfaces gray-white, densely or
sparsely villous; primary lateral segments 3–8-paired; ultimate
segments linear-lanceolate or ovate-lanceolate, sessile, rather
small. Synflorescence terminal, with 1(–3) capitula, not conspicuously
flat-topped. Involucre coryliform, 1–1.4 cm in
diam.; phyllaries in 4 rows, scarious margin dark brown, outer
phyllaries ovate or long ovate-triangular, 3–4(–6) mm, middle
and inner ones narrowly lanceolate or oblanceolate, 5–15 mm,
middle ones abaxially densely villous with fluffy curved long
simple hairs, inner ones glabrous or nearly so abaxially. Ray
florets white or pale red; lamina elliptic or narrowly elliptic, 5–
15 mm, apex 3-denticulate. Achenes ca. 2.5 mm. Corona ca. 1
mm, divided to base. Fl. and fr. Jun–Aug.
Meadows, rocky mountain slopes, floodlands; below 3700 m.
Xinjiang [N India, Kazakhstan, Russia, Uzbekistan].
159. HANDELIA Heimerl, Oesterr. Bot. Z. 71: 215. 1922.
天山蓍属 tian shan shi shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Herbs, perennial; stems 1 to few, thick, with wide pith; indumentum of basifixed hairs. Basal leaves large, petiolate; stem leaves
small, sessile, oblong, 3-pinnatisect. Synflorescences cymose, dense or lax, ± flat-topped. Capitula homogamous, discoid. Involucres
hemispheric; phyllaries in 3 rows, oblong, scarious margins broad. Receptacle conical, paleate; palea narrowly oblong or oblanceolate,
scarious. Florets yellow, bisexual; corolla tubular, apex 5-lobed, gland-dotted. Anther bases obtuse, apical appendage acute.
Style branches claviform-oblong. Achenes small, cuneate, inconspicuously 5-ribbed adaxially, with abaxial rim. Corona absent.
One species: Afghanistan, China, Kazakhstan, Pakistan, Uzbekistan.
1. Handelia trichophylla (Schrenk ex Fischer & C. A. Meyer)
Heimerl, Oesterr. Bot. Z. 71: 215. 1922.
天山蓍 tian shan shi
Achillea trichophylla Schrenk ex Fischer & C. A. Meyer,
Enum. Pl. Nov. 1: 48. 1841.
Herbs, perennial, 30–100 cm tall, with robust taproot; stem
usually solitary, thick, striate, lower part densely dirty white