9. Tribe ANTHEMIDEAE

Tanacetum quercifolium W. W. Smith, Notes Roy. Bot.
Gard. Edinburgh 8: 119. 1913; Chrysanthemum quercifolium
(W. W. Smith) Handel-Mazzetti; Dendranthema quercifolium
(W. W. Smith) Kitamura; Phaeostigma quercifolium (W. W.
Smith) Muldashev.
Subshrubs, 60–150 cm tall. Old branches gray-black;
flowering branches robust, white, powdery-sericeous especially
in upper part and on peduncles. Middle leaves of flowering
branches shortly petiolate; leaf blade narrowly elliptic, lanceolate,
or obovate-oblong, rarely linear, 5–10 × 2–4 cm, rigid,
abaxially white or gray-white, densely and thickly sericeous,
adaxially green, glabrous, margin coarsely dentate, incisedlobed,
or -fid, teeth or lobes 3- or 4-paired, obliquely triangular
or lanceolate, sometimes middle and distal leaves of flowering
branches linear-lanceolate or broadly linear, to 10 cm, margin
entire. Upper leaves progressively smaller. Synflorescence a
dense flat-topped panicle, 4–9 cm in diam.; peduncle robust.
Capitula many. Involucres campanulate, 5–6 mm in diam.;
phyllaries in 4 rows, scarious margin yellow-brown, outer ones
ovate-lanceolate, ca. 2 mm, abaxially sparsely sericeous, middle
and inner ones narrowly elliptic to lanceolate, 3–4 mm, abaxially
glabrous. Florets yellow, with sessile glands; marginal
female florets 11, corolla narrowly tubular, apex minutely 4denticulate;
disk florets many, corolla tubular, apex 5-lobed.
Achenes ca. 1.5 mm. Pappus absent. Fl. and fr. Aug–Oct.
● Forest understories, thickets; 3200–3900 m. Sichuan, Yunnan.
3. Ajania latifolia C. Shih, Bull. Bot. Lab. N. E. Forest. Inst.,
Harbin 6: 12. 1980.
宽叶亚菊 kuan ye ya ju
Chrysanthemum shihchuanum H. Ohashi & Yonekura.
Herbs, perennial, 20–35 cm tall, rhizomes long or short,
procumbent. Flowering and sterile stems many, sometimes
densely clustered, erect, shortly corymbosely branched on
upper part, pubescent, especially on synflorescence branches
and peduncles, densely leafy throughout. Lower leaves withered
by anthesis. Middle and upper leaves sessile or subsessile,
with divided pseudostipules; leaf blade broadly oblanceolate,
obovate, or elliptic, 3–4.5 × 1–2 cm, abaxially gray-white,
densely and thickly appressed pubescent, adaxially green, glabrous,
pinnatifid above middle, base cuneate; lobes 2- or 3paired,
lanceolate or obliquely triangular. Synflorescence a compound
flat-topped panicle, 3–5 cm in diam. Capitula many. Involucres
campanulate, 5–6 mm in diam.; phyllaries in 4 rows,
scarious margin broadly yellow-brown, outer ones ovate or
ovate-lanceolate, 2–2.5 mm, abaxially sparsely pubescent, middle
and inner ones ovate-lanceolate to long ovate, middle ones
sparsely pubescent abaxially, inner ones glabrous abaxially.
Florets yellow; marginal female florets 6, corolla narrowly
tubular, ca. 2.5 mm; disk florets many, corolla tubular, ca. 3
mm, apex 5-lobed. Fl. and fr. Oct.
● Mountain slopes; ca. 3100 m. Sichuan.
4. Ajania variifolia (C. C. Chang) Tzvelev in Schischkin &
Bobrov, Fl. URSS 26: 401. 1961.
异叶亚菊 yi ye ya ju
ANTHEMIDEAE
659
Chrysanthemum variifolium C. C. Chang, Sinensia 5: 161.
1934; Ajania manchurica Poljakov; Dendranthema variifolium
(C. C. Chang) Voroschilov; Phaeostigma variifolium (C. C.
Chang) Muldashev.
Subshrubs, ca. 30 cm tall, old branches with densely leafy
tips. Flowering branches sparsely sericeous or subglabrous.
Middle leaves of flowering branches with petiole 1–2 cm; leaf
blade ovate, 2–3 × 1.5–2.5 cm, abaxially gray-white, densely
sericeous, adaxially green, glabrous, 3–5-pinnatisect; segments
linear or narrowly linear. Lower and upper leaves smaller,
blades 3-sect. Synflorescence a terminal compound flat-topped
panicle. Capitula many. Involucres campanulate, 4–5 mm in
diam.; phyllaries in 4 rows, scarious margin yellow-brown,
outer ones ovate or long ovate, ca. 2 mm, abaxially sparsely
sericeous at base, apex acute, middle and inner ones narrowly
obovate or elliptic, 3–4 mm, apex rounded. Florets yellow,
exterior with sessile glands; marginal female florets 6, corolla
narrowly tubular, ca. 2.5 mm, apex 2- or 3-denticulate; disk
florets many, corolla tubular, ca. 3 mm. Achenes ca. 2 mm. Fl.
and fr. Aug–Sep.
Rocky slopes; 1200–3500 m. Heilongjiang, Hubei, Shaanxi [Korea,
Russia].
The whole plant is used medicinally for relieving spasms.
5. Ajania ramosa (C. C. Chang) C. Shih in C. Shih & G. X. Fu,
Acta Phytotax. Sin. 17(2): 114. 1979.
分枝亚菊 fen zhi ya ju
Chrysanthemum variifolium C. C. Chang var. ramosum C.
C. Chang, Sinensia 5: 163. 1934; C. ramosum (C. C. Chang) H.
Ohashi & Yonekura; Phaeostigma variifolium (C. C. Chang)
Muldashev var. ramosum (C. C. Chang) Muldashev.
Shrubs, 80–150 cm tall. Old branches brownish; flowering
branches of current year sericeous, especially above and on
synflorescence branches. Middle leaves of flowering branches
petiolate; leaf blade elliptic, oblanceolate, or long ovate, 4–5 ×
2–2.5 cm, abaxially white or gray-white, densely and thickly
sericeous, adaxially green, glabrous, pinnatipartite; lobes 3- or
4-paired, narrowly elliptic, lanceolate, or falcate. Proximal and
distal leaves progressively smaller. Capitula compound-corymbose
at apices of branches. Involucres campanulate, 5–6 mm in
diam.; phyllaries in 4 rows, outer ones triangular-ovate, 1.5–2
mm, abaxially sparsely sericeous, scarious margin yellowbrown,
apex rounded; middle and inner ones ovate-oblong or
oblanceolate, 4–4.5 mm. Florets many, corolla yellow, exterior
with sessile glands; marginal female florets 7, corolla narrowly
tubular, ca. 2 mm, apex 4-denticulate; disk florets many, corolla
tubular, ca. 3 mm. Achenes ca. 1.3 mm. Fl. and fr. Aug–Sep. 2n
= 18*.
● Mountain slopes, river valleys; 2900–4600 m. Hubei, Shaanxi,
Sichuan, Xizang.
6. Ajania alabasica H. C. Fu in Ma, Fl. Intramongol. 6: 325.
1982.
内蒙亚菊 nei meng ya ju
Chrysanthemum alabasicum (H. C. Fu) H. Ohashi &
Yonekura.