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9. Tribe ANTHEMIDEAE

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villous or pubescent. Middle stem leaves: petiole ca. 5 mm; leaf<br />

blade orbicular, suborbicular, flabelliform, or broadly cuneate,<br />

0.8–1.5 × 1–1.8 cm, both surfaces gray-white, densely or thickly<br />

appressed pubescent, bipalmatisect or 3–5-palmatisect; primary<br />

lateral segments 1(or 2)-paired; ultimate segments elliptic.<br />

Lower and uppermost stem leaves trisect. Synflorescence a terminal<br />

flat-topped cyme, 2–4 cm in diam. Capitula few, (1–)3–5<br />

or many (to 15). Involucres campanulate, 6–10 mm in diam.;<br />

phyllaries in 4 rows, abaxially densely or sparsely pubescent,<br />

scarious margin dark brown or gray-brown, apex obtuse or<br />

slightly rounded, outer ones lanceolate or linear-lanceolate, 3–4<br />

mm, middle and inner ones broadly lanceolate, narrowly elliptic,<br />

or oblanceolate, 4–5 mm. Florets yellow; marginal female<br />

florets 6–8, corolla narrowly tubular, 3- or 4-denticulate; disk<br />

florets many, corolla tubular. Achenes ca. 1.2 mm. Fl. and fr.<br />

Jul–Sep.<br />

Mountain slopes; 2500–5300 m. Gansu, Nei Mongol, Ningxia,<br />

Qinghai, Sichuan, Xizang, Yunnan [N India].<br />

1<strong>9.</strong> Ajania sericea C. Shih, Bull. Bot. Lab. N. E. Forest. Inst.,<br />

Harbin 6: 14. 1980.<br />

密绒亚菊 mi rong ya ju<br />

Chrysanthemum delavayanum H. Ohashi & Yonekura.<br />

Herbs, perennial, ca. 35 cm tall, with 1 flowering stem and<br />

1 or 2 sterile stems; rhizomes short, with slender fibrous roots.<br />

Flowering stem erect, most parts densely and thickly sericeoustomentose.<br />

Middle stem leaves: petiole 2–4 mm; leaf blade suborbicular<br />

or flabelliform, ca. 0.8 × 1.5 cm, both surfaces graywhite,<br />

densely and thickly tomentose, 3-pinnatisect or subtripalmate-pinnatisect;<br />

primary lateral segments 2-paired; ultimate<br />

segments lanceolate or narrowly elliptic. Lower and upper stem<br />

leaves smaller. Synflorescence a terminal flat-topped panicle,<br />

2–4 cm in diam. Capitula few or moderate in number. Involucres<br />

campanulate, 3–4 mm in diam.; phyllaries in 4 rows,<br />

scarious margin pale brown, outer ones linear or lanceolate, ca.<br />

2.5 mm, abaxially densely and thickly tomentose, middle and<br />

inner ones oblanceolate or narrowly elliptic, ca. 3 mm, middle<br />

ones densely and thickly tomentose abaxially, inner ones glabrous<br />

abaxially. Florets yellow; marginal female florets: corolla<br />

tubular, ca. 1.5 mm, apex 4- or 5-denticulate; disk florets many,<br />

corolla tubular, ca. 2 mm. Fl. May.<br />

● Yunnan (Eryuan).<br />

20. Ajania tripinnatisecta Y. Ling & C. Shih, Bull. Bot. Lab.<br />

N. E. Forest. Inst., Harbin 6: 14. 1980.<br />

多裂亚菊 duo lie ya ju<br />

Chrysanthemum tripinnatisectum (Y. Ling & C. Shih) H.<br />

Ohashi & Yonekura.<br />

Herbs, perennial, ca. 65 cm tall, with slender fibrous roots.<br />

Stems solitary, erect; synflorescence branches short; upper parts<br />

and peduncles appressed pubescent. Middle stem leaves sessile;<br />

leaf blade ovate or elliptic, 3–3.5 × 2–3 cm, abaxially graywhite,<br />

densely appressed pubescent, adaxially green, sparsely<br />

pubescent, 3-pinnatisect; primary lateral segments 3-paired; ultimate<br />

segments lanceolate or elliptic. Upper stem leaves progressively<br />

smaller. Synflorescence a terminal compound flat-<br />

<strong>ANTHEMIDEAE</strong><br />

663<br />

topped panicle. Capitula many. Involucres campanulate, 3–4<br />

mm in diam.; phyllaries in 4 rows, scarious margin yellowbrown,<br />

outer ones elliptic, ca. 2.5 mm, abaxially pubescent,<br />

middle and inner ones elliptic or oblanceolate, 3–4 mm, abaxially<br />

glabrous. Florets yellow, exterior with sessile glands; marginal<br />

female florets 7, corolla narrowly tubular, ca. 2 mm,<br />

acutely 3-denticulate; disk florets many, corolla tubular, ca. 3<br />

mm. Achenes ca. 1 mm. Fl. and fr. Oct.<br />

● Mountain slopes; 3200–3300 m. Sichuan (Hongyuan).<br />

21. Ajania elegantula (W. W. Smith) C. Shih, Bull. Bot. Lab.<br />

N. E. Forest. Inst., Harbin 6: 15. 1980.<br />

云南亚菊 yun nan ya ju<br />

Tanacetum elegantulum W. W. Smith, Notes Roy. Bot.<br />

Gard. Edinburgh 10: 201. 1918; Chrysanthemum elegantulum<br />

(W. W. Smith) S. Y. Hu.<br />

Herbs, perennial, 25–50 cm tall, with small woody rootstock.<br />

Stems many, slender, unbranched, erect or ascending,<br />

thinly silvery gray tomentose. Stem leaves ca. 2 × 1.5 cm, both<br />

surfaces gray tomentulose, 3-pinnatisect; ultimate segments linear,<br />

2–3 × 0.5–0.6 mm. Distal leaves less dissected, those subtending<br />

synflorescence linear, ca. 1 cm. Synflorescence a terminal<br />

flat-topped panicle; peduncles to 1.4 cm, grayish white.<br />

Immature capitula 3–4 mm in diam. Outer phyllaries linear or<br />

lanceolate, 2–3 mm, grayish white tomentulose; middle and<br />

inner ones lanceolate or elliptic, abaxially ± tomentulose, adaxially<br />

straw-colored scarious.<br />

● Yunnan.<br />

Ajania elegantula seems to be known only from the original immature<br />

collections and was regarded as insufficiently known in the<br />

FRPS account of Ajania. However, the elegant, finely dissected leaves<br />

are very distinctive, and it seems justifiable to accept it as a good<br />

species.<br />

22. Ajania nubigena (Wallich ex Candolle) C. Shih in C. Shih<br />

& G. X. Fu, Acta Phytotax. Sin. 17(2): 116. 197<strong>9.</strong><br />

黄花亚菊 huang hua ya ju<br />

Tanacetum nubigenum Wallich ex Candolle, Prodr. 6: 130.<br />

1838; Chrysanthemum nubigenum (Wallich ex Candolle) Handel-Mazzetti;<br />

Dendranthema nubigenum (Wallich ex Candolle)<br />

Kitamura.<br />

Herbs, perennial, ca. 30 cm tall, with slender fibrous roots.<br />

Stems unbranched or branched from base or lower part, densely<br />

pubescent. Middle stem leaves: petiole 3–5 cm; leaf blade<br />

orbicular or broadly ovate, ca. 1 × 1 cm, both surfaces graywhite,<br />

densely pubescent, tripinnatisect or inconspicuously tripinnatisect;<br />

primary lateral segments 2-paired; ultimate segments<br />

linear. Distal stem leaves progressively smaller. Synflorescence<br />

a terminal compound flat-topped panicle, ca. 3 cm in<br />

diam.; synflorescence branches and peduncles white, densely<br />

pubescent. Capitula many. Involucres campanulate, ca. 5 mm in<br />

diam.; phyllaries in 4 rows, abaxially sparsely pubescent, scarious<br />

margin dark brown, apex obtuse or rounded, outer ones<br />

linear-lanceolate, ca. 2.5 mm, middle and inner ones narrowly<br />

elliptic or oblanceolate, 3–4 mm. Florets yellow, exterior with

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