9. Tribe ANTHEMIDEAE

villous or pubescent. Middle stem leaves: petiole ca. 5 mm; leaf
blade orbicular, suborbicular, flabelliform, or broadly cuneate,
0.8–1.5 × 1–1.8 cm, both surfaces gray-white, densely or thickly
appressed pubescent, bipalmatisect or 3–5-palmatisect; primary
lateral segments 1(or 2)-paired; ultimate segments elliptic.
Lower and uppermost stem leaves trisect. Synflorescence a terminal
flat-topped cyme, 2–4 cm in diam. Capitula few, (1–)3–5
or many (to 15). Involucres campanulate, 6–10 mm in diam.;
phyllaries in 4 rows, abaxially densely or sparsely pubescent,
scarious margin dark brown or gray-brown, apex obtuse or
slightly rounded, outer ones lanceolate or linear-lanceolate, 3–4
mm, middle and inner ones broadly lanceolate, narrowly elliptic,
or oblanceolate, 4–5 mm. Florets yellow; marginal female
florets 6–8, corolla narrowly tubular, 3- or 4-denticulate; disk
florets many, corolla tubular. Achenes ca. 1.2 mm. Fl. and fr.
Jul–Sep.
Mountain slopes; 2500–5300 m. Gansu, Nei Mongol, Ningxia,
Qinghai, Sichuan, Xizang, Yunnan [N India].
19. Ajania sericea C. Shih, Bull. Bot. Lab. N. E. Forest. Inst.,
Harbin 6: 14. 1980.
密绒亚菊 mi rong ya ju
Chrysanthemum delavayanum H. Ohashi & Yonekura.
Herbs, perennial, ca. 35 cm tall, with 1 flowering stem and
1 or 2 sterile stems; rhizomes short, with slender fibrous roots.
Flowering stem erect, most parts densely and thickly sericeoustomentose.
Middle stem leaves: petiole 2–4 mm; leaf blade suborbicular
or flabelliform, ca. 0.8 × 1.5 cm, both surfaces graywhite,
densely and thickly tomentose, 3-pinnatisect or subtripalmate-pinnatisect;
primary lateral segments 2-paired; ultimate
segments lanceolate or narrowly elliptic. Lower and upper stem
leaves smaller. Synflorescence a terminal flat-topped panicle,
2–4 cm in diam. Capitula few or moderate in number. Involucres
campanulate, 3–4 mm in diam.; phyllaries in 4 rows,
scarious margin pale brown, outer ones linear or lanceolate, ca.
2.5 mm, abaxially densely and thickly tomentose, middle and
inner ones oblanceolate or narrowly elliptic, ca. 3 mm, middle
ones densely and thickly tomentose abaxially, inner ones glabrous
abaxially. Florets yellow; marginal female florets: corolla
tubular, ca. 1.5 mm, apex 4- or 5-denticulate; disk florets many,
corolla tubular, ca. 2 mm. Fl. May.
● Yunnan (Eryuan).
20. Ajania tripinnatisecta Y. Ling & C. Shih, Bull. Bot. Lab.
N. E. Forest. Inst., Harbin 6: 14. 1980.
多裂亚菊 duo lie ya ju
Chrysanthemum tripinnatisectum (Y. Ling & C. Shih) H.
Ohashi & Yonekura.
Herbs, perennial, ca. 65 cm tall, with slender fibrous roots.
Stems solitary, erect; synflorescence branches short; upper parts
and peduncles appressed pubescent. Middle stem leaves sessile;
leaf blade ovate or elliptic, 3–3.5 × 2–3 cm, abaxially graywhite,
densely appressed pubescent, adaxially green, sparsely
pubescent, 3-pinnatisect; primary lateral segments 3-paired; ultimate
segments lanceolate or elliptic. Upper stem leaves progressively
smaller. Synflorescence a terminal compound flat-
ANTHEMIDEAE
663
topped panicle. Capitula many. Involucres campanulate, 3–4
mm in diam.; phyllaries in 4 rows, scarious margin yellowbrown,
outer ones elliptic, ca. 2.5 mm, abaxially pubescent,
middle and inner ones elliptic or oblanceolate, 3–4 mm, abaxially
glabrous. Florets yellow, exterior with sessile glands; marginal
female florets 7, corolla narrowly tubular, ca. 2 mm,
acutely 3-denticulate; disk florets many, corolla tubular, ca. 3
mm. Achenes ca. 1 mm. Fl. and fr. Oct.
● Mountain slopes; 3200–3300 m. Sichuan (Hongyuan).
21. Ajania elegantula (W. W. Smith) C. Shih, Bull. Bot. Lab.
N. E. Forest. Inst., Harbin 6: 15. 1980.
云南亚菊 yun nan ya ju
Tanacetum elegantulum W. W. Smith, Notes Roy. Bot.
Gard. Edinburgh 10: 201. 1918; Chrysanthemum elegantulum
(W. W. Smith) S. Y. Hu.
Herbs, perennial, 25–50 cm tall, with small woody rootstock.
Stems many, slender, unbranched, erect or ascending,
thinly silvery gray tomentose. Stem leaves ca. 2 × 1.5 cm, both
surfaces gray tomentulose, 3-pinnatisect; ultimate segments linear,
2–3 × 0.5–0.6 mm. Distal leaves less dissected, those subtending
synflorescence linear, ca. 1 cm. Synflorescence a terminal
flat-topped panicle; peduncles to 1.4 cm, grayish white.
Immature capitula 3–4 mm in diam. Outer phyllaries linear or
lanceolate, 2–3 mm, grayish white tomentulose; middle and
inner ones lanceolate or elliptic, abaxially ± tomentulose, adaxially
straw-colored scarious.
● Yunnan.
Ajania elegantula seems to be known only from the original immature
collections and was regarded as insufficiently known in the
FRPS account of Ajania. However, the elegant, finely dissected leaves
are very distinctive, and it seems justifiable to accept it as a good
species.
22. Ajania nubigena (Wallich ex Candolle) C. Shih in C. Shih
& G. X. Fu, Acta Phytotax. Sin. 17(2): 116. 1979.
黄花亚菊 huang hua ya ju
Tanacetum nubigenum Wallich ex Candolle, Prodr. 6: 130.
1838; Chrysanthemum nubigenum (Wallich ex Candolle) Handel-Mazzetti;
Dendranthema nubigenum (Wallich ex Candolle)
Kitamura.
Herbs, perennial, ca. 30 cm tall, with slender fibrous roots.
Stems unbranched or branched from base or lower part, densely
pubescent. Middle stem leaves: petiole 3–5 cm; leaf blade
orbicular or broadly ovate, ca. 1 × 1 cm, both surfaces graywhite,
densely pubescent, tripinnatisect or inconspicuously tripinnatisect;
primary lateral segments 2-paired; ultimate segments
linear. Distal stem leaves progressively smaller. Synflorescence
a terminal compound flat-topped panicle, ca. 3 cm in
diam.; synflorescence branches and peduncles white, densely
pubescent. Capitula many. Involucres campanulate, ca. 5 mm in
diam.; phyllaries in 4 rows, abaxially sparsely pubescent, scarious
margin dark brown, apex obtuse or rounded, outer ones
linear-lanceolate, ca. 2.5 mm, middle and inner ones narrowly
elliptic or oblanceolate, 3–4 mm. Florets yellow, exterior with