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9. Tribe ANTHEMIDEAE

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Roadsides, waste places, steppes, hillsides, forest margins; sea<br />

level to 4200 m. Gansu, N Guizhou, Hebei, Heilongjiang, Jilin, Liaoning,<br />

Nei Mongol, Ningxia, Qinghai, Shaanxi, Shanxi, E and W Sichuan,<br />

Xinjiang, Xizang, W Yunnan [Afghanistan, India, Japan, Kazakhstan,<br />

Korea, Kyrgyzstan, Mongolia, Nepal, Pakistan, Russia, Tajikistan,<br />

Turkmenistan, Uzbekistan; E Europe].<br />

The essential oils from Artemisia sieversiana are used in anti-inflammatory<br />

and hemostatic medicines.<br />

3. Artemisia sericea (Besser) Weber ex Stechmann, Artemis.<br />

16. 1775.<br />

绢毛蒿 juan mao hao<br />

Absinthium sericeum Besser, Bull. Soc. Imp. Naturalistes<br />

Moscou 1: 237. 1829; A. grandiflorum Besser; A. nitens Steven<br />

ex Besser; Artemisia holosericea Ledebour; A. holosericea var.<br />

grandiflora Ledebour; A. holosericea var. parviflora Ledebour;<br />

A. nitens (Steven ex Besser) Steven ex Krascheninnikov; A.<br />

sericea var. gmeliniana Besser; A. sericea var. grandiflora<br />

(Ledebour) Candolle; A. sericea var. ledebouriana Besser; A.<br />

sericea var. nitens (Steven ex Besser) Candolle; A. sericea var.<br />

pallasiana Besser; A. sericea f. parviflora (Ledebour) Pampanini;<br />

A. sericea var. parviflora (Ledebour) Candolle.<br />

Subshrubs or perennial herbs, 40–60(–75) cm tall, with<br />

thick, creeping rhizomes. Stems single to several, usually not<br />

branched, gray sericeous or stems glabrescent. Basal leaves deciduous<br />

by anthesis. Middle stem leaves: petiole 5–8 mm; leaf<br />

blade elliptic or ovate, 3–5 × 2–4 cm, 2-pinnatisect; segments<br />

4(–6) pairs, distal lobes 3-lobulate or subpalmatisect; lobules<br />

linear or linear-lanceolate, 10–17 × 1.5–2 mm, apex acute. Uppermost<br />

leaves and bracts pinnatisect or 3–5-sect. Synflorescence<br />

a ± narrow, racemelike panicle. Capitula shortly pedunculate,<br />

nodding. Involucre hemispheric, (4–)5–6(–9) mm in diam.;<br />

phyllaries with narrow white scarious margin; receptacle hemispheric,<br />

densely pubescent. Marginal female florets 10–14; corolla<br />

narrowly conical or tubular, 2- or 3-toothed. Disk florets<br />

40–80, bisexual; corolla limb puberulent. Achenes ellipsoid or<br />

ellipsoid-ovoid. Fl. and fr. Jul–Oct. 2n = 18.<br />

Forest margins, hills, steppes, canyons, waste areas; 600–1600 m.<br />

Nei Mongol, Ningxia, N Xinjiang [India, Kazakhstan, Mongolia, N<br />

Pakistan, Russia; E Europe].<br />

4. Artemisia aschurbajewii C. Winkler, Trudy Imp. S.-Peterburgsk.<br />

Bot. Sada 11: 332. 1890 [“aschurbajewi”].<br />

褐头蒿 he tou hao<br />

Artemisia sericea (Besser) Weber var. turkestanica C.<br />

Winkler.<br />

Herbs, perennial, ± caespitose, 20–40(–50) cm tall, puberulent.<br />

Stems several, usually not branched. Basal and middle<br />

stem leaves: petiole 8–12 mm; leaf blade suborbicular or orbicular-reniform,<br />

1–2.5(–3.5) × 1–2(–3) cm, 2-pinnatisect; segments<br />

3–5 pairs, 3-lobulate; lobules lanceolate, 5–10 × 2–4<br />

mm, margin serrate, rarely entire, apex obtuse. Uppermost<br />

leaves and bracts 3-lobed; lobes linear or linear-lanceolate. Synflorescence<br />

a racemelike panicle, compact near apex and lax<br />

near base. Capitula shortly pedunculate, nodding. Involucre globose<br />

or subglobose, 5–7 mm in diam.; phyllaries with wide<br />

<strong>ANTHEMIDEAE</strong><br />

679<br />

brown scarious margin; receptacle hemispheric, densely pubescent.<br />

Marginal female florets 10–15; corolla narrowly conical<br />

or tubular, 2- or 3-toothed. Disk florets 40–60, bisexual; corolla<br />

limb puberulent. Achenes ellipsoid. Fl. and fr. Jul–Oct. 2n<br />

= 36.<br />

Deserts or subalpine steppes, rocky hills; 1200–3500 m. NW Gansu,<br />

Qinghai, Xinjiang [Kazakhstan, Kyrgyzstan, Tajikistan].<br />

5. Artemisia absinthium Linnaeus, Sp. Pl. 2: 848. 1753.<br />

中亚苦蒿 zhong ya ku hao<br />

Herbs, perennial, 60–150 cm tall, somewhat woody at<br />

base, gray sericeous or puberulent. Stems 1–3. Basal leaves:<br />

petiole 6–12 cm; leaf blade ovate-elliptic or ovate, 8–12 × 7–9<br />

cm, 2- or 3-pinnatisect; segments 4 or 5 pairs, pinnately lobed;<br />

lobules lanceolate-elliptic or -linear, 8–15 × 2–4(–7) mm, apex<br />

obtuse. Middle stem leaves: petiole 2–6 cm; leaf blade ovate or<br />

elliptic-ovate, 2-pinnatisect; lobules linear-lanceolate, (8–)10–<br />

25 × 2–3(–5) mm. Uppermost leaves 4–6 × 2–4 cm, pinnatisect<br />

or 5-lobed; leaflike bracts 3-lobed or entire; lobes<br />

lanceolate or linear-lanceolate. Synflorescence usually a broad<br />

conical panicle; primary branches straight, ascending or ±<br />

oblique spreading, up to 30 cm and secondary branches up<br />

to 12 cm. Capitula shortly pedunculate, nodding. Involucre globose<br />

or subglobose, 2.5–3.5(–4) mm in diam.; receptacle hemispheric,<br />

densely pubescent. Marginal female florets 15–25; corolla<br />

yellow, obliquely 2-toothed. Disk florets 30–70(–90), bisexual;<br />

corolla yellow. Achenes oblong, 0.8–1 mm, with apical<br />

corona or not. Fl. and fr. Aug–Sep. 2n = 18.<br />

Hillsides, steppes, scrub, forest margins, often in locally moist<br />

situations; 1100–1500 m. Jiangsu, NW and W Xinjiang [Afghanistan,<br />

India, Japan, Kazakhstan, Kyrgyzstan, Pakistan, Russia; N Africa, SW<br />

Asia, Europe, North America].<br />

Artemisia absinthium is used in anti-inflammatory and anthelmintic<br />

medicines.<br />

6. Artemisia frigida Willdenow, Sp. Pl. 3: 1838. 1803.<br />

冷蒿 leng hao<br />

Herbs, perennial, or occasionally subshrubs, (10–)30–<br />

60(–70) cm tall, densely yellowish or pale yellow sericeous,<br />

sometimes stem bases glabrescent. Stems several, woody at<br />

base, shortly branched or not. Lower stem leaves: petiole 5–20<br />

mm; leaf blade oblong or obovate-oblong, 0.8–1.5 × 0.8–1.5<br />

cm, 2(or 3)-pinnatisect; segments (2–)4 pairs; lobules linearlanceolate<br />

or lanceolate. Middle stem leaves: blade oblong or<br />

obovate-oblong, 5–7 × 5–7 mm, 1- or 2-pinnatisect; lobules<br />

lanceolate to linear-lanceolate, 2–3 × 0.5–1.5 mm. Uppermost<br />

leaves and bracts pinnately or 3–5-lobed. Synflorescence a spike<br />

or raceme, sometimes branched to form a panicle. Capitula nodding.<br />

Involucre hemispheric, globose, or ovoid, (2–)2.5–4.5<br />

mm in diam.; receptacle densely white pubescent. Marginal<br />

female florets 8–13; corolla limb yellow or purple. Disk florets<br />

20–30, bisexual. Achenes oblong or ellipsoid-obovate, with<br />

a minute apical corona. Fl. and fr. Jun–Oct. 2n = 18, 36; (n =<br />

9+3b, 18, 18+2b, 36).<br />

Steppes, subalpine meadows, dry hillsides, stable dunes, dry waste

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