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9. Tribe ANTHEMIDEAE

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146. Artemisia oxycephala Kitagawa, Rep. Exped. Manchoukuo<br />

Sect. IV, 4: 93. 1936.<br />

光沙蒿 guang sha hao<br />

Artemisia pubescens Ledebour var. oxycephala (Kitagawa)<br />

Kitagawa.<br />

Subshrubs, 50–80 cm tall, with a thick rootstock. Lowermost<br />

leaves long petiolate; middle stem leaves ± sessile; leaf<br />

blade broadly ovate or suborbicular, 2–5 × 2–3 cm, puberulent<br />

or glabrescent, 2-pinnatisect; segments 2 or 3 pairs; lateral lobes<br />

3-sect; lobules linear, 15–20 × 1.5–2 mm. Uppermost leaves<br />

and leaflike bracts 3–5-sect or entire. Synflorescence a lax, ±<br />

broad panicle. Involucre oblong, 1.5–2.5(–3.5) mm in diam.<br />

Marginal female florets 8–14. Disk florets 3–10, male. Achenes<br />

oblong. Fl. and fr. Aug–Oct.<br />

● Steppes, dry hills, dunes, saline-alkaline soils, lakeshores, forest<br />

steppes; low elevations. N Hebei, W Heilongjiang, W Jilin, W Liaoning,<br />

Nei Mongol, N Shanxi.<br />

147. Artemisia campestris Linnaeus, Sp. Pl. 2: 846. 1753.<br />

荒野蒿 huang ye hao<br />

Oligosporus campestris (Linnaeus) Cassini.<br />

Shrubs, 30–60[–150] cm tall, with a strong woody stock,<br />

much branched, faintly aromatic, pubescent or glabrescent.<br />

Lowermost leaves: petiole 3–5 cm; leaf blade broadly ovate or<br />

ovate, (3–)4–5(–8) × 2.5–5 cm, 2(or 3)-pinnatisect; segments 4<br />

or 5 pairs, 3–5-sect; lobules narrowly linear or narrowly linear-lanceolate,<br />

4–10 × (0.5–)1–1.5[–2] mm, apex mucronulate.<br />

Middle stem leaves sessile, 1- or 2-pinnatisect; segments 3 or 4<br />

pairs. Uppermost leaves and leaflike bracts 3–5-sect or entire.<br />

Synflorescence a lax, broad panicle. Involucre ovoid, 1.5–2.5<br />

mm in diam. Marginal female florets 3–6[–20]. Disk florets 6–<br />

10[–30], bisexual. Achene obovoid. Fl. and fr. Jul–Oct. 2n = 18,<br />

36.<br />

Steppes, waste areas, rocky slopes, dune margins; 300–3100 m. N<br />

Gansu, Taiwan, Xinjiang [Japan, Russia; C Asia, Europe, North<br />

America].<br />

We have not seen material of Artemisia campestris var. glomerata<br />

Pampanini (Nuovo Giorn. Bot. Ital., n.s., 34: 642. 1927), described as<br />

an annual with ± filiform leaf segments.<br />

148. Artemisia marschalliana Sprengel, Syst. Veg. 3: 496.<br />

1826.<br />

中亚旱蒿 zhong ya han hao<br />

Subshrubs or small shrubs, 30–70(–80) cm tall, with a<br />

thick rootstock, gray pubescent or sericeous-pubescent, persistent<br />

or glabrescent. Lower stem leaves: petiole 3–6 cm; leaf<br />

blade oblong-ovate or ovate, (3–)4–8(–10) × 2–5(–6) cm, 2(or<br />

3)-pinnatisect; segments 4–6 pairs, 3–5-lobuled. Middle and upper<br />

stem leaves petiolate; leaf blade 1- or 2-pinnatisect; segments<br />

3 or 4(or 5) pairs; lobules linear or linear-lanceolate, 6–<br />

15 × (0.5–)1–1.5 mm, apex mucronulate. Leaflike bracts 3–5sect<br />

or entire. Synflorescence a broad to slender, erect or ascending,<br />

conical panicle, sometimes with lateral branches to<br />

form compound panicle; branches short, mostly less than 1 cm.<br />

Capitula contiguous along ultimate branches, ± sessile, erect.<br />

<strong>ANTHEMIDEAE</strong><br />

725<br />

Involucre ovoid or broadly ovoid, 1.5–2.5(–3) mm in diam.<br />

Marginal female florets 3–8. Disk florets 5–15, male. Achenes<br />

ovoid. Fl. and fr. Jul–Oct.<br />

Steppes, forest steppes, wastelands, rocky slopes, hills; 500–2200<br />

m. N Xinjiang [Kazakhstan, Russia; Europe].<br />

1a. Stem, branches, leaves, and phyllaries<br />

gray pubescent, later glabrescent;<br />

synflorescence an ascending,<br />

slender conical panicle ............... 148a. var. marschalliana<br />

1b. Stem, branches, leaves, and phyllaries<br />

persistently sericeous-pubescent;<br />

synflorescence a broad, compound<br />

panicle ............................................. 148b. var. sericophylla<br />

148a. Artemisia marschalliana var. marschalliana<br />

中亚旱蒿(原变种) zhong ya han hao (yuan bian zhong)<br />

Artemisia inodora M. Bieberstein, Fl. Taur.-Caucas. 2:<br />

295. 1808, not Miller (1768), nor Willdenow (1809); A. campestris<br />

Linnaeus var. gmeliniana Besser; A. campestris var.<br />

marschalliana (Sprengel) Poljakov; A. campestris var. steveniana<br />

Besser; A. tomentella Trautvetter var. subglabra Krascheninnikov;<br />

Oligosporus marschallianus (Sprengel) Lessing.<br />

Stems with few branches above base; vegetative parts and<br />

phyllaries gray pubescent, later glabrescent. Synflorescence a<br />

slender, ascending conical panicle.<br />

Steppes, forest steppes, wastelands, rocky slopes, hills; 500–2200<br />

m. N Xinjiang [Kazakhstan, Russia; Europe].<br />

148b. Artemisia marschalliana var. sericophylla (Ruprecht)<br />

Y. R. Ling, Bull. Bot. Res., Harbin 8(4): 48. 1988.<br />

绢毛旱蒿 juan mao han hao<br />

Artemisia sericophylla Ruprecht, Beitr. Pflanzenk. Russ.<br />

Reiches 2: 41. 1845; A. campestris var. sericophylla (Ruprecht)<br />

Poljakov.<br />

Stem, branches, leaves, and phyllaries persistently sericeous-pubescent.<br />

Stems branching to form a broad compound<br />

panicle.<br />

Steppes, forest steppes, wastelands, rocky slopes, hills; 500–2200<br />

m. N Xinjiang [Kazakhstan, Russia; Europe].<br />

14<strong>9.</strong> Artemisia saposhnikovii Krascheninnikov ex Poljakov,<br />

Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17:<br />

412. 1955.<br />

昆仑沙蒿 kun lun sha hao<br />

Subshrubs, 30–40 cm tall, with a thick rootstock and capitate<br />

branches. Lower and middle stem leaves: petiole 3–15 mm;<br />

leaf blade ovate, 1–3 × 0.5–2.5 cm, yellowish pubescent or glabrescent,<br />

1(or 2)-pinnatisect; segments 2 pairs, linear, straight<br />

or falcate, 3–8(–15) × 0.5–1 mm, margin deeply serrate, apex<br />

obtuse and mucronulate. Uppermost leaves and leaflike bracts<br />

pinnatisect, 3-sect, or entire. Synflorescence a narrow panicle.<br />

Capitula contiguous along ultimate branches, nodding. Involucre<br />

oblong-ovoid or ellipsoid, 2–2.5 mm in diam. Marginal<br />

female florets 4 or 5. Disk florets 4–6, male. Achenes oblong or<br />

oblong-obovoid. Fl. and fr. Aug–Oct. 2n = 36.

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