9. Tribe ANTHEMIDEAE

770
and 2 lateral thick whitish ribs, sometimes also 1 or 2 thin
abaxial ribs, abaxially and apically with 2 distinct resin sacs.
Corona with broad obtuse teeth. Fl. and fr. Jun–Aug.
Sunny mountain slopes, grasslands of river valleys; 700–2600 m.
Heilongjiang, Xinjiang (Altay Shan, Tian Shan) [Kazakhstan, Mongolia,
Russia; SW Asia].
2. Tripleurospermum homogamum G. X. Fu ex Y. Ling & C.
Shih, Bull. Bot. Lab. N. E. Forest. Inst., Harbin 6: 9. 1980.
无舌三肋果 wu she san lei guo
Herbs, perennial, ca. 40 cm tall, with short rhizomes; stems
unbranched, striate, conspicuously pubescent below capitula.
Basal leaves: petiole short, dilated basally; leaf blade oblanceolate-oblong,
4–6 × 0.8–1.5 cm, pinnatisect to 2-pinnatisect,
both surfaces glabrous or very sparsely pubescent; ultimate segments
linear, sessile. Distal stem leaves linear, entire. Capitula
solitary, terminal, homogamous, discoid; peduncle to 10 cm. Involucres
hemispheric, ca. 1 cm in diam.; phyllaries in 4 rows,
abaxially glabrous or sparsely pubescent, scarious margin dark
brown, outer and middle phyllaries lanceolate to oblong-lanceolate,
ca. 4 mm, inner ones spatulate-oblanceolate, ca. 3.5 mm.
Ray florets absent. Disk florets yellow, 5-lobed, lobes each with
a resin sac. Achenes pale brown, 3-angled, ca. 2 mm, smooth,
with 1 adaxial and 2 lateral thick whitish ribs, sometimes 1 or 2
thin abaxial ribs, abaxially and apically with 2 distinct resin
sacs. Corona ca. 0.2 mm, 3-lobed.
● Alpine forest margins; ca. 2500 m. N Xinjiang (Burqin).
3. Tripleurospermum tetragonospermum (F. Schmidt)
Pobedimova, Bot. Mater. Gerb. Bot. Inst. Komarova Akad.
Nauk S.S.S.R. 21: 346. 1961.
东北三肋果 dong bei san lei guo
Chamaemelum tetragonospermum F. Schmidt, Reis.
Amur-Land., Bot. 148. 1868; Matricaria tetragonosperma (F.
Schmidt) H. Hara & Kitamura.
Herbs, annual, 40–50 cm tall; stems usually branched from
base, striate, upper parts sparsely pubescent, lower part glabrous.
Lower and middle stem leaves sessile; leaf blade oblanceolate-oblong
or oblong, 5–15 × 2–5 cm, 2- or 3-pinnatisect,
both surfaces glabrous; ultimate segments linear-filiform. Distal
stem leaves similar, smaller. Capitula solitary, terminal, several
per plant, 2–2.5 cm in diam. Involucres hemispheric, 1.2–1.5
cm in diam.; phyllaries in 4 rows, abaxially glabrous, scarious
margin narrow, brown, outer ones ovate-oblong, apex obtuse,
middle ones narrowly oblong, apex rounded, inner ones lanceolate-oblong,
apex obtuse. Ray florets white; lamina ca. 6.5 mm.
Disk florets yellow, 5-lobed. Achenes brownish, triquetrous, ca.
1.5 mm, tuberculate, with 1 adaxial and 2 lateral thick ribs,
abaxially and apically with 2 distinct resin sacs. Corona ca. 0.2
mm, entire. Fl. and fr. Jun–Aug.
Roadsides, sandy places by rivers; ca. 300 m. Heilongjiang, Liaoning
[Japan, Russia].
4. Tripleurospermum inodorum (Linnaeus) Schultz Bipontinus,
Tanaceteen, 32. 1844.
ANTHEMIDEAE
新疆三肋果 xin jiang san lei guo
Matricaria inodora Linnaeus, Fl. Suec., ed. 2, 297. 1755;
Chamaemelum inodorum (Linnaeus) Visiani; Chamomilla inodora
(Linnaeus) K. Koch; Chrysanthemum inodorum (Linnaeus)
Linnaeus; C. maritimum (Linnaeus) Cavanilles var. inodorum
(Linnaeus) Becherer; Dibothrospermum agreste Knaf,
nom. illeg. superfl.; M. maritima Linnaeus subsp. inodora (Linnaeus)
Soó; M. maritima var. inodora (Linnaeus) Soó; Pyrethrum
inodorum (Linnaeus) Moench; Rhytidospermum inodorum
(Linnaeus) Schultz Bipontinus; Tripleurospermum maritimum
(Linnaeus) W. D. J. Koch subsp. inodorum (Linnaeus)
Applequist.
Herbs, annual or biennial; stems 30–70 cm tall, erect, striate,
glabrous, corymbosely branched in upper 1/2. Middle stem
leaves sessile; leaf blade ovate-oblong or oblong, 2–4 × 1–2.5
cm, both surfaces glabrous, 2-pinnatisect; ultimate segments
narrowly linear. Distal stem leaves similar, gradually smaller.
Capitula several to many, in an apical corymb, 2–3 cm in diam.,
pedunculate. Involucres hemispheric, 7–10 mm in diam.; phyllaries
in 3 or 4 rows, margin narrowly white scarious, outer
ones lanceolate, middle and inner ones oblong to oblanceolate.
Ray florets white; lamina ca. 1 cm. Disk florets yellow, 5lobed;
lobes each with a resin sac. Achenes brown, triquetrous,
ca. 2 mm, wrinkled, with 1 adaxial and 2 lateral thick whitish
ribs, abaxially and apically with 2 distinct resin sacs. Corona
short, subentire. Fl. and fr. Sep. 2n = 36.
River valleys; ca. 1100 m. Jiangsu, Jilin, Liaoning, N Xinjiang
(Tacheng) [Kazakhstan, Russia, Uzbekistan; Europe].
5. Tripleurospermum limosum (Maximowicz) Pobedimova,
Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 21:
352. 1961.
三肋果 san lei guo
Chamaemelum limosum Maximowicz, Mém. Acad. Imp.
Sci. St.-Pétersbourg Divers Savans 9: 156. 1859; Matricaria
limosa (Maximowicz) Kudô; M. maritima Linnaeus subsp. limosa
(Maximowicz) Kitamura.
Herbs, annual or biennial; stems 10–35 cm tall, striate,
glabrous, unbranched or branched from base. Basal leaves withered
by anthesis. Lower and middle stem leaves sessile; leaf
blade oblanceolate-oblong or oblong, 5.5–9.5 × 2.5–3 cm, both
surfaces glabrous, tripinnatisect; ultimate segments filiform.
Distal stem leaves gradually smaller, similar to proximal and
middle leaves. Capitula few to many, in an apical corymb, 1–
1.5 cm in diam., pedunculate. Involucres hemispheric; phyllaries
in 2 or 3 rows, margin white or brownish scarious, outer
ones broadly lanceolate, inner ones oblong. Ray florets white;
lamina 4(–6) mm. Disk florets yellow, 5-lobed; lobes each with
a resin sac. Achenes brown, triquetrous, ca. 2.5 mm, wrinkled,
with 1 adaxial and 2 lateral whitish ribs, apically and abaxially
with 2 distinct resin sacs. Corona ca. 0.5 mm, 3-lobed. Fl. and
fr. Jun–Aug.
Sandy places by rivers and lakes, meadows, dry sandy mountain
slopes. Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol [Japan, Kazakhstan,
Korea, Mongolia, Russia, Uzbekistan].