9. Tribe ANTHEMIDEAE

ded; ultimate segments oblong to linear, to 1.5 mm wide. Upper
stem leaves reduced, subsessile, 3–5-lobed. Synflorescence a
terminal flat-topped panicle. Capitula 20–60. Involucres 2.5–3
mm in diam.; phyllaries herbaceous, abaxially subglabrous,
scarious margin broad, light or brown, apex broadened, middle
ANTHEMIDEAE
667
and inner phyllaries 2–3 × as long as outer, to 4 mm. Florets
15–25, corolla tubular; marginal female florets few; disk florets:
corolla 1.5–2 mm. Achenes 1.2–1.5 mm.
Stony slopes; above 3000 m. Ningxia, Xizang [Kyrgyzstan, Tajikistan
(Pamirs)].
148. AJANIOPSIS C. Shih, Acta Phytotax. Sin. 16(2): 86. 1978.
画笔菊属 hua bi ju shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Herbs, annual. Leaves alternate, biternately pinnatisect, 2-pinnatisect, or trisect. Synflorescences cymose; capitula many or few,
in ± flat-topped panicles, rather small, heterogamous, disciform. Involucres obovoid, 4–5 mm in diam.; phyllaries in 2 rows.
Receptacle convex, epaleate. Marginal florets in 1 row, female; corolla pitcher-shaped, tapering above; limb yellow, apex minutely 2toothed,
apically pilose with straight erect hairs. Disk florets many, bisexual; corolla tubular, apex 5-lobed. Anther bases obtuse;
apical appendage subtriangular. Style branches truncate at apex. Achenes obovoid, 5- or 6-ribbed. Pappus absent.
● One species: China.
1. Ajaniopsis penicilliformis C. Shih, Acta Phytotax. Sin.
16(2): 87. 1978.
画笔菊 hua bi ju
Plants 5–10 cm tall. Stems few branched from base,
branches prostrate in lower part, upper part ascending, white
villous. Middle stem leaves: petiole ca. 0.5 mm; leaf blade
broadly cuneate or suborbicular, 0.5–0.7 × ca. 1 cm, both
surfaces densely villous, biternately pinnatisect or 2-pinnatisect;
primary lateral segments 1-paired; ultimate segments linear or
broadly linear, apex acute. Proximal and distal leaves small.
Synflorescence a flat-topped terminal panicle. Capitula many or
few. Involucres obovoid, 4–5 mm in diam.; phyllaries in 2
rows, abaxially densely villous, scarious margin dark, outer
ones elliptic, 3–3.5 mm, inner ones spatulate or oblanceolate,
ca. 3 mm. Florets yellow, apically pilose with erect straight
hairs; marginal female florets 7 or 8, corolla ascidiform, ca. 2
mm, tapering above; disk florets 12, corolla tubular, ca. 2.5
mm. Achenes obovoid, ca. 1.8 mm, 5- or 6-ribbed. Pappus absent.
Fl. and fr. Sep.
● Steep slopes; 4600–5000 m. Xizang (Nanmulin).
149. BRACHANTHEMUM Candolle, Prodr. 6: 44. 1838.
短舌菊属 duan she ju shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Subshrubs, ± procumbent, woody at base, pubescent with basifixed, medifixed, or stellate hairs. Leaves alternate or subopposite,
few lobed. Synflorescences cymose; capitula solitary or in ± flat-topped cymes or panicles, heterogamous, radiate, rarely
homogamous and discoid. Involucres campanulate, hemispheric, or obconical; phyllaries in 4 or 5 rows, rigidly herbaceous, scarious
margin white or brown. Receptacle flat or convex to conical, epaleate. Marginal florets female, fertile, rarely absent; lamina yellow,
rarely white. Disk florets yellow, bisexual; corolla tubular, apex 5-lobed. Anther bases obtuse; apical appendage lanceolate. Style
branches linear, apex truncate. Achenes terete, 5-striate. Corona absent.
Ten species: C Asia, China, Mongolia; six species (one endemic) in China.
1a. Capitula homogamous, ray florets absent; phyllaries abaxially prominently gland-dotted .......................................... 6. B. gobicum
1b. Capitula heterogamous, ray florets present; phyllaries abaxially not or only obscurely gland-dotted.
2a. Plants with appressed stellate hairs ..................................................................................................................... 5. B. pulvinatum
2b. Plants with simple or medifixed hairs.
3a. Most leaves palmately 3(–5)-lobed.
4a. Leaves sessile or subsessile, leaf blade obliquely elliptic or suborbicular, ca. 0.6 × 0.5 cm;
achenes ca. 2.8 mm ............................................................................................................................... 1. B. mongolicum
4b. Leaves rather long petiolate, leaf blade almost orbicular, to 1.5(–2) × ca. 1.5 cm including
petiole; achenes 1.8–2.2 mm ............................................................................................................... 2. B. fruticulosum
3b. Most leaves pinnately 4–7-lobed, uppermost leaves sometimes palmately 3-lobed.
5a. Capitula (1–)3–15(–20), 2–4 crowded at apex, remainder on long divergent peduncles ............................. 3. B. titovii
5b. Capitula 1(–5) on long erect peduncles ................................................................................................ 4. B. kirghisorum