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9. Tribe ANTHEMIDEAE

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766<br />

few, similar to basal leaves, sessile. Synflorescence an irregular<br />

flat-topped panicle. Capitula 5–12; peduncle 6–8 cm, not thickened.<br />

Involucre campanulate, 4–7 mm in diam.; phyllaries in 4<br />

rows, scarious margin narrow, outer phyllaries lanceolate, ca.<br />

1.5 mm, inner ones narrowly linear-elliptic, 3–5 mm. Ray florets<br />

yellow. Achenes ca. 2 mm. Corona 0.2–0.4 mm, lobed or<br />

serrate. Fl. and fr. Jun–Aug.<br />

Rocky slopes, moist places on mountain slopes; 1100–2100 m.<br />

Xinjiang [Kazakhstan, Russia].<br />

6. Tanacetum scopulorum (Krascheninnikov) Tzvelev in<br />

Schischkin & Bobrov, Fl. URSS 26: 342. 1961.<br />

岩菊蒿 yan ju hao<br />

Pyrethrum scopulorum Krascheninnikov, Bot. Mater.<br />

Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 9: 164. 1946;<br />

Lepidolopsis scopulorum (Krascheninnikov) Poljakov.<br />

Herbs, perennial, to 35 cm tall, with branched rhizomes;<br />

stems solitary or clustered, upper parts shortly corymbosely<br />

branched, densely or sparsely pilose with T-shaped and simple<br />

hairs. Basal leaves: petioles to 2 cm; leaf blade narrowly elliptic<br />

or elliptic, 4–8 × 1–2 cm, inconspicuously 3-pinnatisect, both<br />

surfaces green or pale gray-green, pilose; primary lateral<br />

segments 8–10-paired; ultimate segments ovate-lanceolate or<br />

obliquely triangular. Stem leaves few, sessile. Synflorescence a<br />

flat-topped cyme. Capitula 3–6; peduncle 1–8 cm. Involucre<br />

campanulate, 7–10 mm in diam.; phyllaries in 4 rows, abaxially<br />

villous or pubescent, outer phyllaries lanceolate, ca. 2.5 mm,<br />

apex white scarious-ampliate, middle and inner ones elliptic to<br />

linear-elliptic, 3–5 mm. Ray florets yellow, apex 3- or 4-denticulate.<br />

Achenes 2–2.3 mm. Corona 0.2–0.3 mm, margin serrate.<br />

Fl. and fr. Jul–Aug.<br />

Mountain slopes; ca. 700 m. Xinjiang (Altay Shan) [Kazakhstan].<br />

7. Tanacetum cinerariifolium (Treviranus) Schultz Bipontinus,<br />

Tanaceteen, 58. 1844.<br />

除虫菊 chu chong ju<br />

Pyrethrum cinerariifolium Treviranus, Index Sem. Hort.<br />

Bot. Wratislav. App. 2: 2. 1820; Chrysanthemum cinerariifolium<br />

(Treviranus) Visiani.<br />

Herbs, perennial; stems 17–60 cm tall, solitary or fascicled,<br />

unbranched or branched from base, pubescent with appressed<br />

T-shaped or furcate hairs. Basal leaves: petiole 10–20<br />

cm; leaf blade ovate or elliptic, 1.5–4 × 1–2 cm, both surfaces<br />

silvery grayish, appressed pubescent with T-shaped and furcate<br />

hairs, 2-pinnatisect; primary segments 3–5-paired; ultimate<br />

segments linear or oblong-ovate, margin entire or few dentate.<br />

Lower and middle leaves similar, large, shortly petiolate.<br />

Capitula apically solitary or 3–10 in apical lax corymb. Involucre<br />

cup-shaped, 1.2–1.5 cm in diam.; phyllaries in 4 rows, abaxially<br />

pubescent especially in outer ones, outer ones lanceolate,<br />

ca. 4 mm, middle and inner ones lanceolate to broadly linear,<br />

5–6 mm, margin narrowly white scarious. Ray florets white;<br />

lamina 1.2–1.5 cm, apex truncate or emarginate. Achenes 2.5–<br />

3.5 mm. Corona 0.8–1.5 mm, margin shallowly lobed. Fl. and<br />

fr. May–Aug. 2n = 18*, 24, 27, 29, 31, 34, 36.<br />

<strong>ANTHEMIDEAE</strong><br />

Widely cultivated. Anhui, Guizhou, Hebei, Liaoning, Zhejiang<br />

[native to SE Europe; now widely cultivated].<br />

Tanacetum cinerariifolium is widely cultivated, mainly in tropical<br />

upland regions, as a source of pyrethrins, which are extracted from the<br />

dried capitula and used as insecticides. Pyrethroids are important insecticides<br />

obtained by the chemical modification of pyrethrins.<br />

8. Tanacetum coccineum (Willdenow) Grierson, Notes Roy.<br />

Bot. Gard. Edinburgh 33: 262. 1974.<br />

红花除虫菊 hong hua chu chong ju<br />

Chrysanthemum coccineum Willdenow, Sp. Pl. 3: 2144.<br />

1803; C. marschallii Ascherson; Pyrethrum coccineum (Willdenow)<br />

Voroschilov.<br />

Herbs, perennial; stems 25–50 cm tall, erect, solitary,<br />

unbranched or few branched from base, subglabrous. Basal<br />

leaves: petiole 2–10 cm; leaf blade ovate or narrowly elliptic,<br />

4–8 × 2.5–4 cm, 2-pinnatisect, both surfaces sparsely pubescent<br />

or glabrous; primary lateral segments 5-paired; ultimate segments<br />

triangular, elliptic, or narrowly elliptic. Stem leaves<br />

similar, sessile or shortly petiolate; leaves below capitula small,<br />

pinnatifid or entire. Capitula solitary, terminal. Involucre cupshaped,<br />

1–1.5 cm in diam.; phyllaries in 4 rows, scarious margin<br />

brown, outer phyllaries lanceolate, ca. 3.5 mm, abaxially<br />

glabrous or sparsely pubescent, middle and inner ones narrowly<br />

elliptic or linear-oblanceolate, ca. 5 mm, abaxially glabrous.<br />

Ray florets red; lamina narrowly elliptic, ca. 1.6 cm, apex 2- or<br />

3-denticulate. Achenes ca. 2.5 mm. Corona 0.1–0.2 mm, margin<br />

shallowly obtusely lobed. Fl. and fr. May–Oct. 2n = 36.<br />

Cultivated. Anhui, Hebei [native to SW Asia (Caucasus)].<br />

Tanacetum coccineum is used as an insecticide in the same way as<br />

T. cinerariifolium and is also grown as an ornamental.<br />

<strong>9.</strong> Tanacetum petraeum (C. Shih) K. Bremer & Humphries,<br />

Bull. Nat. Hist. Mus. London, Bot. 23: 103. 1993 [“petrareum”].<br />

岩匹菊 yan pi ju<br />

Pyrethrum petraeum C. Shih, Bull. Bot. Lab. N. E. Forest.<br />

Inst., Harbin 6: 10. 1980 [“petrareum”].<br />

Subshrubs, ca. 35 cm tall; old branches gray; young<br />

branches turning purple-red, sparsely pubescent or glabrous.<br />

Lower leaves: petiole ca. 1.5 cm; leaf blade ovate, obovate, or<br />

oblanceolate, 2–4 × 1.4–2.5 cm, 2-pinnatipartite, both surfaces<br />

glabrous; primary lateral lobes 2- or 3-paired; ultimate lateral<br />

lobes 1-paired, obliquely triangular. Distal leaves pinnatifid,<br />

small. Synflorescence an irregular flat-topped cyme with 1–3<br />

capitula. Involucre coryliform, 15–20 mm in diam.; phyllaries<br />

in 4 rows, abaxially densely pubescent to glabrescent or glabrous,<br />

scarious margin dark brown or brown, outer phyllaries<br />

narrowly oblanceolate-elliptic, ca. 6 mm, middle and inner ones<br />

lanceolate-triangular or elliptic, 7–8 mm, middle ones glabrescent,<br />

inner ones glabrous. Ray florets pink; lamina narrowly elliptic,<br />

ca. 2 cm, apex 2-denticulate. Achenes ca. 2 mm. Corona<br />

0.1–0.2 mm, divided 1/2 length or to base. Fl. and fr. Aug.<br />

● On rocks of mountain slopes; 1800–2600 m. Xinjiang (Fuyun).

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