9. Tribe ANTHEMIDEAE

754
ANTHEMIDEAE
1a. Rosette or cushion plants with concealed stems, to 7 cm tall, usually less; capitula in very dense sessile clusters.
2a. Rosette plants with a single central cluster of capitula.
3a. Leaves 2-pinnatisect ....................................................................................................................................... 8. H. glomerata
3b. Leaves 3–7-pinnatisect ........................................................................................................................ 9. H. syncalathiformis
2b. Cushion plants with several distinct clusters of capitula.
4a. Leaves spatulate, apex 3–6-lobed ................................................................................................................ 10. H. gossypina
4b. Leaves orbicular or flabelliform, biternate-palmately divided ...................................................................... 11. H. kennedyi
1b. Plants with obvious stems, 7–50 cm tall; capitula in pedunculate panicles.
5a. Capitula fascicled-corymbose at apices of branches or stem; involucres campanulate, glossy, yellowish or
straw-colored, phyllaries in 4 or 5 rows, scarious margins pale brownish or white.
6a. Perennial herbs, often with a solitary stem ....................................................................................................... 3. H. delavayi
6b. Subshrubs.
7a. Leaves 2-pinnatisect, abaxially green or deep green, glabrous; florets ca. 3.2 mm .............................. 2. H. desmantha
7b. Leaves pinnatipartite, abaxially gray-white, densely appressed pubescent; florets ca. 2 mm ............ 1. H. kaschgarica
5b. Capitula corymbose or glomerulate; involucres cuneate or cuneate-campanulate, matt, phyllaries in 3 or
4 rows, scarious margins dark brown.
8a. Leaves 3-pinnatisect; capitula many .......................................................................................................... 4. H. yunnanensis
8b. Leaves 2-pinnatisect; capitula 4–12.
9a. Plants to 30 cm tall; leaves narrowly elliptic or ovate, 2–4 cm, ultimate segments narrowly elliptic ........ 5. H. herderi
9b. Plants to 12 cm tall; leaves ovate, ultimate segments linear.
10a. Stem leaves sessile ......................................................................................................................... 6. H. senecionis
10b. Stem leaves petiolate, petiole 5–10 mm ........................................................................................ 7. H. tomentosa
1. Hippolytia kaschgarica (Krascheninnikov) Poljakov, Bot.
Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 18: 290.
1957.
贺兰山女蒿 he lan shan nü hao
Tanacetum kaschgaricum Krascheninnikov, Trudy Bot.
Inst. Akad. Nauk S.S.S.R., Ser. 1, Fl. Sist. Vyssh. Rast. 1: 175.
1933; Chrysanthemum alashanense (Y. Ling) Y. Ling; Hippolytia
alashanensis (Y. Ling) C. Shih; Poljakovia alashanensis
(Y. Ling) Grubov & Filatova; P. kaschgarica (Krascheninnikov)
Grubov & Filatova; T. alashanense Y. Ling.
Subshrubs or small shrubs, 25–50 cm tall; old branches
dark gray or gray-brown; branchlets of current year somewhat
gray-white, indumentum dense and appressed powdery. Leaves
in basal rosettes and scattered along stems; petiole 3–8 mm; leaf
blade narrowly elliptic, lanceolate, or elliptic, 1.5–2 × 0.5–1
cm, pinnatipartite, abaxially gray-white, densely appressed pubescent,
adaxially green, glabrescent; lateral lobes 2–5-paired,
all lobe margins entire or 1- or 2-toothed on one side; leaves
below synflorescences sometimes simple, entire. Synflorescence
a dense terminal flat-topped cyme; capitula 3–10, peduncle
0.5–1.5 cm, pubescent with appressed white powdery hairs.
Involucre campanulate, 4–5 mm in diam.; phyllaries in 4 rows,
rigidly herbaceous, glossy, yellowish or straw-colored, scarious
margin brown or white, outer phyllaries lanceolate, ca. 1.8 mm,
middle and inner ones oblanceolate, ca. 3 mm. Florets yellow,
ca. 2 mm. Achenes ca. 1.5 mm. Fl. and fr. Aug–Oct.
● Mountain slopes, fissures of rocks, grasslands, desert steppes;
1900–2300 m. Gansu, Nei Mongol, Ningxia, Xinjiang.
Tzvelev (Fl. URSS 26: 412. 1961) suggested that this species
would have to be removed from Hippolytia. Bremer and Humphries
(Bull. Nat. Hist. Mus. London, Bot. 23: 106. 1993) also regarded this
species as anomalous, and it was subsequently transferred by Grubov
and Filatova to the new genus Poljakovia (Novosti Sist. Vyssh. Rast. 33:
227. 2001).
Filatova (Rast. Tsentral. Azii 14a: 30–31. 2007) treated Hippolytia
alashanensis and H. kaschgarica as distinct species in Poljakovia: P.
alashanensis with lower cauline leaves and leaves of sterile shoots pinnatisect,
lateral segments linear, apex shortly cuspidate; and P. kaschgarica
with lower cauline leaves and leaves of sterile shoots pinnatifid,
lateral segments oblong-lanceolate, entire, apex subobtuse.
2. Hippolytia desmantha C. Shih, Acta Phytotax. Sin. 17(4):
63. 1979.
束伞女蒿 shu san nü hao
Subshrubs, 10–15 cm tall; old branches gray; young
branches greenish or brown-green, pubescent with appressed
powdery hairs. Leaves with petioles ca. 5 mm; leaf blade ovate,
elliptic, obliquely elliptic, or narrowly flabelliform, 0.5–2 × 1–
1.5 cm, both surfaces green, glabrous, 2-pinnatisect; primary
lateral segments 2- or 3-paired; ultimate segments linear, narrowly
elliptic, or lanceolate; leaves below synflorescence sometimes
pinnatisect. Capitula 3–5, fascicled-corymbose at apices
of branches, pedunculate; peduncle slender, ca. 1.5 cm, pubescent
with appressed powdery hairs. Involucres campanulate, 4–
5 mm in diam.; phyllaries in 4 rows, rigidly herbaceous, glossy,
yellow-white or straw-colored, margin brownish, outer ones
triangular-ovate, 1.5–2 mm, middle and inner ones narrowly elliptic
or oblanceolate, ca. 4 mm. Florets yellow, ca. 3.2 mm.
Achenes ca. 2 mm. Fl. and fr. Aug.
● Meadows, rock outcrops on valley slopes; 3800–3900 m. Qinghai
(Yushu).
3. Hippolytia delavayi (Franchet ex W. W. Smith) C. Shih,
Acta Phytotax. Sin. 17(4): 65. 1979.
川滇女蒿 chuan dian nü hao
Tanacetum delavayi Franchet ex W. W. Smith, Notes Roy.
Bot. Gard. Edinburgh 8: 345. 1915; Chrysanthemum bulbosum
(Handel-Mazzetti) Handel-Mazzetti; C. delavayi (Franchet ex
W. W. Smith) Handel-Mazzetti; T. bulbosum Handel-Mazzetti.