9. Tribe ANTHEMIDEAE

ANTHEMIDEAE
1a. Leaves 4–5 cm; blade oblong, pinnatisect or pinnatipartite; ray florets female; achenes pilose, with distinct corona.
2a. Leaves arachnoid lanate on both surfaces; capitula 1.2–2 cm wide; achenes ca. 3 mm, pilose ......................... 1. A. tomentosa
2b. Leaves glabrous on both surfaces; capitula to 1 cm wide; achenes ca. 1.5 mm, glabrescent .............................. 2. A. stoliczkae
1b. Leaves 0.8–2 cm; blade spatulate or cuneate, palmately 3–5-lobed or -parted; ray florets sometimes sterile;
achenes mostly glabrous (pilose in A. lasiocarpa), with reduced corona.
3a. Achenes pilose ...................................................................................................................................................... 8. A. lasiocarpa
3b. Achenes glabrous.
4a. Plants glabrous or tomentose only below involucre and on peduncle; achene 2.5–4 mm.
5a. Capitula 2.5–3.5 cm in diam., peduncles glabrous; phyllaries 6–7 mm, glabrous, apex cuspidate;
achenes 2.5–4 mm ................................................................................................................................. 3. A. tridactylites
5b. Capitula 1.5–2.5 cm in diam., peduncles sparsely hairy; phyllaries 3.5–4.5 mm, basally hairy,
apex ± obtuse; achenes 2.5–3.5 mm ............................................................................................................. 4. A. glabra
4b. Plants grayish white lanate; achenes 1.2–2 mm.
6a. Plants rather sparsely lanate; capitula ca. 2 cm in diam.; leaves 8–12 mm ................................................ 5. A. huegelii
6b. Plants densely lanate, hairs thick.
7a. Capitula 3.5–4 cm in diam.; leaves 10–15 mm, distinctly 3–5-lobed ................................................... 6. A. vestita
7b. Capitula 1.5–2 cm in diam.; leaves 4–8 mm, obscurely 3-lobed ............................................................. 7. A. nivea
1. Allardia tomentosa Decaisne in Jacquemont, Voy. Inde
4(Bot.): 87. 1841.
羽裂扁毛菊 yu lie bian mao ju
Waldheimia tomentosa (Decaisne) Regel.
Herbs, perennial, 10–15 cm tall, with procumbent muchbranched
rhizomes. Stems many, laxly caespitose, branched or
not, bases with remains of brown scarious leaf sheaths. Petioles
enlarged at base; leaf blade oblong to linear-oblong, 4–5 × 0.2–
1.5 cm, 2-pinnatipartite, both surfaces white arachnoid lanate,
primary lateral pinnae 6–9-paired, distal pinnae gradually
smaller; ultimate pinna lanceolate to ovate, apex long mucronulate.
Capitulum solitary, pedunculate. Involucre hemispheric,
17–20 mm in diam.; phyllaries in 3 or 4 rows, abaxially lanate,
scarious margin broad, dark brown, outer ones lanceolate, short,
apex acute, inner ones linear-oblong, subscarious. Ray florets
10–20, white or pale pink; lamina linear-oblong, 6–11 mm.
Disk florets yellow, 5-lobed. Achenes narrowly oblong, ca. 4.5
mm, 6–8-ribbed; ribs pale red-brown in upper part, sparsely
villous in upper 1/2, with sessile glands; corona scales many,
ca. 6 mm, brownish at apex. Fl. and fr. Aug–Sep.
Talus on mountain slopes; 4200–5200 m. W Xizang (Burang) [Afghanistan,
India, Kazakhstan, Pakistan, Uzbekistan].
2. Allardia stoliczkae C. B. Clarke, Compos. Ind. 145. 1876.
光叶扁毛菊 guang ye bian mao ju
Waldheimia korolkowii Regel & Schmalhausen; W. stoliczkae
(C. B. Clarke) Ostenfeld.
Herbs, perennial, ca. 15 cm tall, with procumbent, manybranched
rhizomes; stems glabrous. Leaf blades 4–5 × 1(–1.5)
cm, glabrous, 1- or 2-pinnatisect. Capitulum solitary, terminal,
pedunculate. Phyllaries glabrous, scarious margin dark. Ray floret
lamina white or pale pink, ca. 15 mm. Achenes sparsely pilose
or glabrous.
Rocky slopes, stony riverbeds; [3000–5000 m]. Xinjiang, W Xizang
[Afghanistan, India, Kazakhstan, Pakistan, Uzbekistan].
No Chinese specimens were seen. The account is based on Fl.
URSS (26: 268. 1961) and Rast. Tsentral. Azii (14a: 23. 2007).
749
3. Allardia tridactylites (Karelin & Kirilov) Schultz Bipontinus,
Pollichia 20–21: 442. 1863.
三指扁毛菊 san zhi bian mao ju
Waldheimia tridactylites Karelin & Kirilov, Bull. Soc.
Imp. Naturalistes Moscou 15: 126. 1842.
Herbs, perennial, to 6 cm, with woody, procumbent,
much-branched rhizome; stems many; entire plant glabrous.
Leaves in dense rosettes, sessile; leaf blade cuneate, 1–2 × 0.5–
1 cm, glabrous, base cuneate-attenuate, apex 3(–5)-lobed or
-parted. Capitulum solitary, terminal, 2.5–3.5 cm in diam., sessile.
Involucre hemispheric, 1.5–2 cm in diam.; phyllaries in 3
or 4 rows, abaxially glabrous, scarious margin broad, dark
brown, outer phyllaries ovate-oblong to oblong, ca. 7 mm, apex
cuspidate, inner ones oblong, ca. 8 mm. Ray florets 8–15; lamina
pinkish white, pink, or purple-red, elliptic-oblong, 7–11 ×
2.5–4 mm, apex 2- or 3-denticulate. Disk florets: corolla yellow,
4–5.5 mm, apex 5-lobed. Achenes 2.5–4 × 0.6–0.8 mm,
somewhat curved, 5-ribbed, glabrous, with sessile glands. Corona
scales many, bristlelike, usually tinged reddish or brownish,
6.5–8 mm. Fl. Jul–Sep.
Floodlands, talus on mountain slopes; 3000–4000 m. Xinjiang,
Xizang [Kazakhstan, Mongolia, Russia].
4. Allardia glabra Decaisne in Jacquemont, Voy. Inde 4(Bot.):
88. 1841.
扁毛菊 bian mao ju
Waldheimia glabra (Decaisne) Regel; W. tridactylites
Karelin & Kirilov subsp. glabra (Decaisne) Podlech.
Herbs, perennial, to 4 cm, with woody, procumbent,
much-branched rhizomes; stems many; entire plant grayish or
grayish green, loosely tomentose to pubescent with long simple
hairs. Leaves in dense rosettes, sessile; leaf blade cuneate,
1–1.5 × 0.5–0.8 cm, both surfaces glabrous, base cuneate-attenuate,
apex 3(–5)-lobed or -parted. Capitulum solitary, terminal,
2.5–3.5 cm in diam., sessile. Involucre hemispheric, 1–1.5 cm
in diam.; phyllaries in 3 or 4 rows, abaxially hairy at base, scarious
margin broad, dark brown, apex ± obtuse, outer phyllaries
ovate-oblong to oblong, ca. 7 mm, inner ones oblong, ca. 8 mm.