9. Tribe ANTHEMIDEAE

772
ANTHEMIDEAE
Three species: Mediterranean region, introduced elsewhere as weeds or cultivated as an ornamental or as vegetables; three species (all introduced)
in China.
1a. Ray achenes narrowly 2-winged; leaves irregularly dentate or rarely pinnatilobed ...................................................... 3. G. segetum
1b. Ray achenes widely 3-winged; leaves bipinnately divided.
2a. Adaxial wings of ray achenes projected into an apical rostrum ............................................................................. 1. G. carinata
2b. Wings of ray achenes not projected into rostrum .................................................................................................. 2. G. coronaria
1. Glebionis carinata (Schousboe) Tzvelev, Bot. Zhurn. 84(7):
117. 1999.
蒿子杆 hao zi gan
Chrysanthemum carinatum Schousboe, Iagttag. Vextrig.
Marokko, 198. 1800; Ismelia carinata (Schousboe) Schultz Bipontinus;
I. versicolor Cassini, nom. illeg. superfl.
Herbs, annual, 20–70 cm tall, glabrous or nearly so; stems
erect, usually branched above middle. Basal leaves withered
by anthesis; lower and middle stem leaves sessile; leaf blade
obovate or narrowly elliptic, 8–10 × 3–4 cm, 2-pinnatipartite;
primary lateral lobes 3–8-paired; ultimate lobes lanceolate,
obliquely triangular or linear. Synflorescence a terminal cyme.
Capitula (1 or)2–8, long pedunculate. Involucres cupular, 1.5–
2.5 cm in diam.; phyllaries in 4 rows, inner ones ca. 1 cm.
Ray achenes broadly 3-winged, wings adaxially projected into
an apical rostrum. Disk achenes with 2 elevated ribs. Corona
absent.
Cultivated as a vegetable; 500–2200 m. Anhui, Guizhou, Hainan,
Hebei, Heilongjiang, Hubei, Hunan, Jiangsu, Jilin, Liaoning, Nei Mongol,
Qinghai, Shandong, Xinjiang, Yunnan, Zhejiang [native to NW
Africa (Morocco); introduced elsewhere].
2. Glebionis coronaria (Linnaeus) Cassini ex Spach, Hist. Nat.
Vég. 10: 181. 1841.
茼蒿 tong hao
Chrysanthemum coronarium Linnaeus, Sp. Pl. 2: 890.
1753, nom. cons.; C. roxburghii Desfontaines; Glebionis roxburghii
(Desfontaines) Tzvelev; Matricaria coronaria (Linnaeus)
Desrousseaux; Pinardia coronaria (Linnaeus) Lessing;
Pyrethrum indicum Sims (1813), not (Linnaeus) Cassini (1826);
Xantophtalmum coronarium (Linnaeus) P. D. Sell.
Herbs, annual, glabrous or almost so; stems erect, 70 cm
tall, unbranched or few branched above middle. Basal leaves
withered by anthesis; lower and middle stem leaves sessile;
leaf blade narrowly elliptic or elliptic-lanceolate in outline, 8–
10 × 3–4 cm, 1 or 2(or 3)-pinnatisect or pinnatifid, primary seg-
ments 2–4-paired, ultimate lobes ovate or linear; upper stem
leaves gradually smaller. Capitula terminal, solitary or few, not
forming conspicuous panicle; peduncle 15–20 cm. Involucres
cup-shaped, 1.5–3 cm in diam.; phyllaries in 4 rows, inner ca. 1
cm, apex widened, scarious. Ray achenes narrowly 3-winged.
Disk achenes 1- or 2-ribbed. Fl. and fr. Sep. 2n = 18*
Widely cultivated as a vegetable. Anhui, Fujian, Guangdong,
Guangxi, Guizhou, Hainan, Hebei, Hunan, Jilin, Shandong, Zhejiang
[native to the Mediterranean region].
Glebionis coronaria is grown both for its bright yellow florets
and the edible young shoots. Various authors include Chrysanthemum
coronarium var. spatiosum within this species, but it is better placed
within the following species, G. segetum.
3. Glebionis segetum (Linnaeus) Fourreau, Ann. Soc. Linn.
Lyon, sér. 2, 17: 90. 1869.
南茼蒿 nan tong hao
Chrysanthemum segetum Linnaeus, Sp. Pl. 2: 889. 1753;
C. coronarium Linnaeus f. spatiosum (L. H. Bailey) Kitamura;
C. coronarium var. spatiosum L. H. Bailey; C. spatiosum (L. H.
Bailey) L. H. Bailey; C. umbrosum Willdenow; Matricaria
segetum (Linnaeus) Schrenk; Pyrethrum segetum (Linnaeus)
Moench; P. umbrosum (Willdenow) Boissier; Xantophtalmum
segetum (Linnaeus) Schultz Bipontinus.
Herbs, annual, 20–60 cm tall, glabrous or nearly so; stems
erect, fleshy. Basal and lower stem leaves sessile; leaf blade
elliptic, obovate-lanceolate, or ovate-elliptic, margin irregularly
largely dentate, rarely pinnatilobed. Upper stem leaves gradually
smaller. Synflorescence ill-defined, not obviously flattopped;
capitula solitary or few at apices of branches; peduncle
ca. 5 cm. Involucres cupular, 1–2 cm in diam.; phyllaries in 4
rows, inner ones scarious, apex enlarged. Ray achenes narrowly
2-winged. Disk achenes with 10 ribs.
Cultivated as a vegetable. Anhui, Beijing, Fujian, Guangdong,
Guizhou, Hainan, Hubei, Hunan, Jiangsu, Jiangxi, Yunnan, Zhejiang
[native to the Mediterranean region].
173. LEUCANTHEMUM Miller, Gard. Dict. Abr., ed. 4, [769]. 1754.
滨菊属 bin ju shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Herbs, perennial; indumentum absent or of simple basifixed hairs. Leaves alternate, margin entire, serrate, or pinnatifid. Synflorescences
cymose, lax, ± flat-topped, sometimes reduced to single capitulum. Capitula pedunculate, heterogamous, radiate. Involucres
coryliform; phyllaries in 3 or 4 rows, scarious margins white or brown. Receptacle convex, sometimes conical, epaleate.
Marginal florets in 1 row, female, fertile; lamina white or pink, rarely yellow. Disk florets many, yellow, bisexual; corolla tubular,
apex 5-lobed. Anther bases obtuse; apical appendage ovate-lanceolate. Style branches linear, apex truncate. Achenes 10-ribbed, ribs
projected or not to apical rim. Corona or adaxial auricle present or not.