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9. Tribe ANTHEMIDEAE

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748<br />

topped cyme; capitula 1–10(–15). Involucres hemispheric, 4–5<br />

mm in diam.; phyllaries in 3 rows, ovate to broadly ovate, margin<br />

scarious, apex rounded. Florets yellow. Marginal female<br />

florets 6; corolla ascidiform, tapering above, minutely 2–4-denticulate,<br />

exterior with sessile glands. Disk florets many, tubular,<br />

ca. 2 mm, 5-lobed. Achenes dark, obovate or elliptic, slightly<br />

<strong>ANTHEMIDEAE</strong><br />

compressed, glabrous, 2-striate adaxially. Fl. and fr. Jun–Sep.<br />

2n = 16, 18.<br />

Grasslands on mountain slopes; 1500–2600 m. Hebei, Heilongjiang,<br />

Jilin, Liaoning, Nei Mongol, Shanxi [Japan, Korea, Mongolia,<br />

Russia (Siberia)].<br />

155. NEOPALLASIA Poljakov, Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk<br />

S.S.S.R. 17: 42<strong>9.</strong> 1955.<br />

栉叶蒿属 zhi ye hao shu<br />

Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert<br />

Herbs, annual or perennial. Leaves alternate, pectinate to pinnatisect. Synflorescence of short axillary spikes grouped into a<br />

slender, leafy panicle. Capitula rather small, heterogamous, disciform. Involucre ovoid or broadly ovoid; phyllaries in 3 or 4 rows,<br />

scarious margins broad. Receptacle narrowly conical, epaleate. Marginal florets usually 3 or 4, female, fertile; corolla narrowly<br />

tubular, apex not toothed. Disk florets of two kinds; outer bisexual, inner completely male with reduced ovaries; all central florets<br />

with corolla apex 5-lobed. Anther bases obtuse; apical appendage rhomboid-acuminate. Style branches linear. Achenes in a row<br />

around base of receptacle, ellipsoid, somewhat compressed or triquetrous, brown, striate. Corona absent.<br />

One species: China, Kazakhstan, Mongolia, Russia.<br />

Molecular data (Watson et al., BMC Evolutionary Biology; http://www.biomedcentral.com/1471-2148/2/17/; accessed 10 Apr 2011) indicate<br />

that Neopallasia is closely related to Seriphidium s.s. and could be included with that genus within a more broadly conceived Artemisia.<br />

1. Neopallasia pectinata (Pallas) Poljakov, Bot. Mater. Gerb.<br />

Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 430. 1955.<br />

栉叶蒿 zhi ye hao<br />

Artemisia pectinata Pallas, Reise Russ. Reich. 3: 755.<br />

1776; A. pectinata var. yunnanensis Pampanini; A. yunnanensis<br />

(Pampanini) Krascheninnikov (1958), not Jeffrey ex Diels (Jun<br />

1912), nor H. Léveillé (Nov 1912); Neopallasia tibetica Y. R.<br />

Ling; N. yunnanensis (Pampanini) Y. R. Ling.<br />

Herbs, 12–40 cm tall; stems erect, unbranched or branched<br />

from base, purplish, ± densely white sericeous. Lower and middle<br />

stem leaves sessile; leaf blade oblong-elliptic, 1.5–3 ×<br />

0.5–1 cm, both surfaces green, glabrous, sometimes with sessile<br />

glands, pectinately pinnatisect; segments linear-subulate, somewhat<br />

rigid. Upper stem leaves to below synflorescence similar,<br />

but smaller. Synflorescence of short axillary spikes grouped into<br />

a slender, leafy panicle. Capitula many, rather small, sessile.<br />

Involucres ovoid or broadly ovoid, 2.5–3 mm in diam.; phyllaries<br />

in 3 or 4 rows, abaxially glabrous, scarious margin<br />

broad, outer ones somewhat short, inner ones rather narrow.<br />

Marginal female florets 3 or 4, fertile, narrowly tubular, without<br />

apical teeth. Disk florets tubular, 5-lobed, of two kinds:<br />

outer bisexual, inner completely male with reduced ovaries.<br />

Achenes brown, elliptic, 1.2–1.5 mm, somewhat compressed.<br />

Deserts, gravelly places of river valleys, wastelands; 1300–3400<br />

m. Gansu, Hebei, Heilongjiang, Jilin, Liaoning, Nei Mongol, Ningxia,<br />

Qinghai, Shanxi, Sichuan, Xinjiang, Xizang [Kazakhstan, Mongolia,<br />

Russia].<br />

Neopallasia pectinata is a highly variable species, and the Chinese<br />

entities N. tibetica and N. yunnanensis seem best treated as synonymous<br />

with it.<br />

156. ALLARDIA Decaisne in Jacquemont, Voy. Inde 4(Bot.): 87. 1841.<br />

扁毛菊属 bian mao ju shu<br />

Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert<br />

Waldheimia Karelin & Kirilov.<br />

Herbs, perennial, glabrous to densely tomentose. Leaves alternate, closely spaced, sometimes rosulate, spatulate or cuneate,<br />

apex 3–5-lobed or -parted, or oblong, pinnatisect or pinnatipartite. Capitula solitary, pedunculate, heterogamous, radiate. Involucre<br />

hemispheric; phyllaries in 3 or 4 rows, scarious margins dark brown. Receptacle convex, epaleate. Marginal florets in 1 row, female<br />

or neuter, usually sterile; lamina pink, white, or bluish violet. Disk florets many, yellow or bluish violet, bisexual, fertile; corolla<br />

tubular, apex 5-lobed. Anther bases obtuse, apical appendage ovate-lanceolate. Style branches linear, apex truncate. Achenes somewhat<br />

curved, faintly 5–10-ribbed, generally with sessile glands, sometimes pilose. Corona of 20–50 bristlelike scales, with flat and<br />

palmate base, apex brownish, sometimes wider.<br />

Eight species: Afghanistan, China, India, Kazakhstan, Mongolia, Nepal, Pakistan, Russia, Tajikistan, Uzbekistan; eight species (one endemic) in<br />

China.<br />

The exact date of publication of Allardia is uncertain, but such evidence as there is (Zain-ul-Abidin, Biologia 4: 212–214. 1958) suggests that it<br />

was in 1841, thus predating Waldheimia, which was published in 1842.

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