9. Tribe ANTHEMIDEAE

ANTHEMIDEAE
2. Artemisia sect. Abrotanum (Duhamel du Monceau) Besser, Bull. Soc. Imp. Naturalistes Moscou 1: 222. 1829.
艾蒿组 ai hao zu
Abrotanum Duhamel du Monceau, Traité Arbres Arbust. 1: 19. 1755.
Herbs, annual or perennial, rarely subshrubs, pubescent, tomentose, or glabrous, rarely arachnoid, without glandular or viscid
hairs (sessile glands often present). Leaves 2–4-pinnatisect; lobules pectinate, filiform, narrowly linear, or lanceolate, less than 1(–1.5)
mm wide. Capitula hemispheric, globose, or subglobose, rarely ovoid. Phyllary margins scarious, midvein green or sometimes colored.
Receptacle glabrous. Marginal florets (1–)3–20; corolla narrowly tubular or rarely narrowly conical, 2- or 3(or 4)-toothed or
without teeth. Disk florets (7–)10–80, bisexual, ovaries well developed; style ca. as long as or longer than corolla, branches divergent,
recurved at apex.
About 65 species: N Hemisphere, mostly in Asia and Europe, a few species extending into N Africa, Oceania, and the Americas; 30 species
(seven endemic) in China.
Most species contain mainly monoterpenoids and also a few sesquiterpenoids. This section has been confused with Artemisia sect. Artemisia.
1a. Leaves pinnatisect, with ultimate lobules filiform, linear-filiform, filiform-clavate, narrowly lanceolate, or
linear-lanceolate, (5–)10+ × 0.5–1(–1.5) mm, apex acute.
2a. Herbs annual, biennial, or short-lived perennial; rootstock a simple taproot, sometimes ± fusiform.
3a. Herbs biennial or short-lived perennial; capitula obconical, 4–5 mm in diam. ................................................ 52. A. fukudo
3b. Herbs annual; capitula ovoid or subglobose, 2–3 mm in diam.
4a. Middle stem leaves pinnatisect, segments 2(or 3) pairs; plants pubescent, ± glabrescent ................. 53. A. przewalskii
4b. Middle stem leaves 2- or 3-pinnatisect, segments (2 or)3 or 4 pairs; plants glabrous or nearly so.
5a. Stem branches short or absent; middle stem leaves (1 or)2-pinnatisect, segments (2 or)3 or
4 pairs; capitula 2–10 grouped, in narrow panicles ........................................................................... 46. A. palustris
5b. Stem branches divaricate; middle stem leaves 2- or 3-pinnatisect, segments 4 or 5 pairs;
capitula 2–5 grouped, in broad lax panicles ......................................................................................... 47. A. aurata
2b. Herbs perennial, subshrubs, or shrubs; rootstock often branched, ± woody.
6a. Middle stem leaves 2- or 3-pinnatisect, lobules narrowly linear, densely silvery gray or yellowish
sericeous-tomentose; corolla limb puberulent ............................................................................................... 49. A. austriaca
6b. Middle stem leaves 1- or 2-pinnatisect, lobules linear, filiform, or filiform-clavate, pubescent or
arachnoid pubescent, sometimes glabrescent later, or densely white tomentose abaxially; corolla
limb glabrous.
7a. Stem much branched; middle stem leaves 2–4 cm, 2(or 3)-pinnatisect; abaxial leaf surface and
phyllaries densely white tomentose .................................................................................................... 50. A. brachyloba
7b. Stem shortly capitellate branched or not; middle stem leaves 0.8–2.5(–3) cm, 1- or 2-pinnatisect;
leaves and phyllaries puberulent or leaves glabrescent.
8a. Stem terete; lobules of middle stem leaves filiform, apex acute ....................................................... 44. A. adamsii
8b. Stem 4-ribbed; lobules of middle stem leaves filiform-clavate or linear-filiform, apex
obtuse ........................................................................................................................................... 45. A. dalai-lamae
1b. Leaves pinnatipartite or pinnatisect, with ultimate lobules pectinate, serrate, or smoothly obtusely lacerate,
less than 5(–8) × 1.5(–2) mm, or lobules minute but then lobes 1–2 mm wide with obtuse apex.
9a. Herbs annual or biennial; stem solitary, more than 20 cm; rootstock a simple taproot, sometimes ± fusiform.
10a. Leaves ± succulent, middle stem leaves 1- or 2-pinnatisect, segments 1 or 2 pairs, with obtuse apex,
entire or 1–3-crenate or cleft .................................................................................................................... 51. A. succulenta
10b. Leaves thinly papery, middle stem leaves 2- or 3-pinnatisect, segments 2 or more pairs, lobes or
lobules acute or obtuse apically.
11a. Basal leaves many, forming rosettes, segments more than 20 pairs; phyllaries with brown or
purple-brown scarious margins; marginal female florets 3–8 ............................................................... 41. A. hedinii
11b. Basal leaves not forming distinct rosettes, segments 3–8(–10) pairs; marginal female
florets 10–12.
12a. Involucre 5–7 mm in diam.; phyllaries puberulent; disk florets 80–150; middle
stem leaf segments 3 or 4 pairs, lobes serrate ................................................................. 38. A. shangnanensis
12b. Involucre 1.5–5(–7) mm in diam.; phyllaries glabrescent; disk florets 10–40;
middle stem leaf segments 4–10 pairs, lobes pectinate.
13a. Lower stem leaf petioles 2–6 cm; leaf blades minutely and lanceolately
pectinate; synflorescence narrow and spikelike ..................................................... 42. A. tournefortiana
13b. Lower stem leaf petioles less than 2 cm; leaf blades pectinate; synflorescence
broader, often a conical, compound panicle.
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