9. Tribe ANTHEMIDEAE

● Mountain slopes; 2000–2200(–2600) m. Xinjiang.
18. Tanacetum artemisioides Schultz Bipontinus ex J. D.
Hooker, Fl. Brit. India 3: 318. 1881.
艾状菊蒿 ai zhuang ju hao
Herbs, perennial, to 60 cm tall, from woody rootstock;
stems several, adpressed hairy, glabrescent. Basal and lower
stem leaves petiolate. Middle stem leaves subsessile to sessile;
leaf blade oblong, 1–2.5(–3) × 0.8–1.5(–2) cm, ± palmately 1-
or 2-pinnatisect; ultimate segments oblong, 2–3 mm, obtuse.
Synflorescence a ± lax flat-topped cyme. Capitula 3–10; peduncle
short to moderately long. Involucre ± cupuliform, ca. 5 mm
in diam.; phyllaries in 3 rows, pubescent, margin membranous;
outer phyllaries lanceolate, ca. 3 mm, apex acute; middle and
inner ones oblanceolate to narrowly obovate, 4–5 mm, apex obtuse.
Receptacle convex, glabrous. All florets yellow; corolla
tubular, 3–4 mm, 5-toothed. Achenes light brown.
2400–2700 m. W Xizang [?India, Pakistan].
The type gatherings of this and the following species, Tanacetum
falconeri, were both collected from the border area between Xizang and
ANTHEMIDEAE
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Pakistan. The two species are therefore likely to be found within China.
The generic position of these species with homogamous capitula needs
more detailed investigation.
19. Tanacetum falconeri J. D. Hooker, Fl. Brit. India 3: 320.
1881.
西藏菊蒿 xi zang ju hao
Herbs, perennial or rarely annual, 40–60(–70) cm tall,
from woody rootstock; stems sulcate. Basal and lower stem
leaves petiolate; leaf blade linear-oblong, 1–9(–13) cm including
petiole, 2-pinnatisect, both surfaces silky tomentose;
ultimate segments linear, acute. Synflorescence a flat-topped
cyme. Capitula 6–10(–12), homogamous, discoid; peduncle
1.5–3 cm. Involucre 2–3(–4) mm; phyllaries in 4 rows, abaxially
softly tomentose, lanate, or pubescent, margin membranous.
Receptacle convex, rarely flat. All florets yellow; corolla
tubular, 2–4 mm, 5-toothed. Achenes light brown, 1–1.5(–3)
mm. Corona glandular, entire.
Rocky talus, around lakes, grassy ridges, valley plains; 2000–4000
m. W Xizang [N India (Himachal Pradesh), Pakistan].
170. TRIPLEUROSPERMUM Schultz Bipontinus, Tanaceteen, 31. 1844.
三肋果属 san lei guo shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Chamaemelum Visiani (1845), not Miller (1754); Dibothrospermum Knaf; Rhytidospermum Schultz Bipontinus.
Herbs, annual or perennial; indumentum absent or of simple, basifixed hairs. Leaves 2- or 3-pinnatisect. Synflorescences cymose;
capitula solitary or in ± flat-topped panicles, pedunculate, heterogamous or homogamous, radiate or discoid. Involucres hemispheric;
phyllaries in 2–4 rows, scarious margins often narrow, white or pale to dark brown. Receptacle convex to conical, epaleate.
Marginal florets in 1 row, female, lamina white or rarely pale pink. Disk florets bisexual; corolla tubular, apex 5-lobed; lobes each
usually with resin sac. Anther bases obtuse, apical appendage ovate-triangular or oblong. Style branches with apex truncate. Achenes
3-angled, often wrinkled or tuberculate, with 1 adaxial and 2 lateral, and sometimes 1 or 2 abaxial, white ribs, abaxially and apically
with (1 or)2(–5) distinct resin sacs. Corona present, sometimes forming an auricle, or with a few scales, sometimes absent.
Thirty-eight species: Europe and temperate Asia, a few species in N Africa and North America; five species (one endemic) in China.
1a. Plants perennial, with rhizome; phyllaries with dark brown scarious margins.
2a. Capitula heterogamous, radiate ............................................................................................................................ 1. T. ambiguum
2b. Capitula homogamous, discoid ........................................................................................................................ 2. T. homogamum
1b. Plants annual or biennial, without rhizome; phyllaries with white or paler brown scarious margins.
3a. Phyllaries with narrow pale brown scarious margin; achenes tuberculate ............................................. 3. T. tetragonospermum
3b. Phyllaries with white or pale brown scarious margin; achenes wrinkled.
4a. Ray floret lamina ca. 10 × 3.5 mm; corona subentire; leaf blade 2–4 × 1–2.5 cm ......................................... 4. T. inodorum
4b. Ray floret lamina ca. 4 × 1.5 mm; corona 3-toothed; leaf blade 5.5–9.5 × 2.5–3 cm ...................................... 5. T. limosum
1. Tripleurospermum ambiguum (Ledebour) Franchet &
Savatier, Enum. Pl. Jap. 1: 236. 1875.
褐苞三肋果 he bao san lei guo
Pyrethrum ambiguum Ledebour, Fl. Altaic. 4: 118. 1833;
Chamaemelum ambiguum (Ledebour) Boissier; Matricaria
ambigua (Ledebour) Krylov.
Herbs, perennial, 15–50 cm tall, rhizomatous; stems erect,
unbranched, striate, scattered pilose below capitula. Basal leaves
with petiole base dilated; leaf blade lanceolate-oblong or oblong,
3–7(–10) × 1–2 cm, 2-pinnatisect, both surfaces glabrous;
primary lateral segments many paired; ultimate segments linear,
sessile, small. Capitula solitary, terminal, 3–4 cm in diam. Involucre
hemispheric, 1–1.5 cm in diam.; phyllaries in 3 rows,
abaxially glabrous, scarious margin narrow, dark brown, outer
phyllaries lanceolate, 6–6.5 mm, middle ones oblong, inner
ones oblanceolate. Ray florets white; lamina ca. 2 × 4.5 mm.
Disk florets yellow, 5-lobed; lobes each with red apical resin
sac. Achenes deep brown, ca. 2 mm, tuberculate, with 1 adaxial