9. Tribe ANTHEMIDEAE

lanate, upper part glabrescent. Basal leaves petiolate; leaf blade
oblong, 5–30 × 2–12 cm, 3-pinnatisect, densely pilose; ultimate
segments subfiliform, sessile, smaller. Synflorescence lax or
dense, ± flat-topped. Involucres hemispheric, ca. 5 mm in
diam.; phyllaries in 3 rows, oblong, scarious margin broad,
adaxially sparsely villous. Florets many, yellow, tubular, 5-
ANTHEMIDEAE
753
lobed, gland-dotted. Achenes yellow-brown, cuneate, 1–1.5 ×
0.3–0.5 mm, with abaxial rim. Corona absent.
Xinjiang [Afghanistan, Kazakhstan, Pakistan, Uzbekistan].
The description above is based on specimens from Kazakhstan
and Uzbekistan and on the account in Fl. URSS (26: 125. 1961).
160. PSEUDOHANDELIA Tzvelev in Schischkin & Bobrov, Fl. URSS 26: 878. 1961.
拟天山蓍属 ni tian shan shi shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Herbs, biennial to perennial, dying after flowering; stems 1–4, thick with wide pith; indumentum of basifixed hairs, arachnoid.
Leaves alternate, 2- or 3-pinnatisect. Capitula many in dense pseudoumbel, long pedunculate, homogamous, discoid. Involucre hemispheric;
phyllaries in 2 or 3 rows, margins scarious, pale; innermost phyllaries resembling receptacle scales. Receptacle hemispheric
to conical, epaleate or with a few marginal palea. Florets yellow, bisexual, fertile, narrowly tubular with abruptly expanded campanulate
limb, 5-lobed; tube glandular papillate. Anthers without basal appendages; apical appendage ovate-lanceolate, apex obtuse.
Style branches linear, apex truncate. Achenes narrowly cylindric, curved, circular in cross section, inconspicuously 4- or 5-ribbed,
glandular-papillate. Pappus absent.
One species: Afghanistan, China, Kazakhstan, Tajikistan, Turkmenistan; SW Asia (Iran).
1. Pseudohandelia umbellifera (Boissier) Tzvelev in Schischkin
& Bobrov, Fl. URSS 26: 363. 1961.
拟天山蓍 ni tian shan shi
Tanacetum umbelliferum Boissier, Diagn. Pl. Orient., ser.
2, 3: 30. 1856; Chrysanthemum floccosum Kitamura; C. trichophyllum
(Regel & Schmalhausen) Kuntze; C. umbelliferum
(Boissier) Hoffmann; Lepidolopsis umbellifera (Boissier) Poljakov;
Pyrethrum trichophyllum (Regel & Schmalhausen)
Bornmüller & Sintenis (1900–1901), not Grisebach (1846); P.
umbelliferum (Boissier) Boissier; T. trichophyllum Regel &
Schmalhausen.
Herbs, biennial or short-lived perennial, dying after flowering,
20–100 cm tall, roots long and thick; stems solitary or
few, robust, unbranched below synflorescence, base covered
with persistent leaf bases; most parts densely arachnoid hairy.
Leaf blade to 30 × 5–6 cm, bipinnate, ultimate segments linear,
to 0.6 mm wide, with cartilaginous cusp. Synflorescence umbel-like.
Capitula many, long pedunculate. Involucres hemispheric,
3–5 × 6–9 mm; outer phyllaries oblong, margin broadly
scarious, erose; innermost phyllaries much narrower. Florets
many; corolla yellow, with linear tube and abruptly expanded
campanulate limb, 5-lobed, strongly glandular papillate.
Achenes cylindric, 1.8–2.4 × ca. 0.3 mm, obscurely 4- or 5ribbed,
tuberculate, glandular. Corona absent.
Stony slopes, sands, gravel beds, etc. Xinjiang [Afghanistan, Kazakhstan,
Tajikistan, Turkmenistan; SW Asia (Iran)].
No material from China was seen by the present authors. The
description is based primarily on the account in Fl. URSS.
161. HIPPOLYTIA Poljakov, Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk
S.S.S.R. 18. 288. 1957.
女蒿属 nü hao shu
Shi Zhu (石铸 Shih Chu); Christopher J. Humphries, Michael G. Gilbert
Herbs, perennial, sometimes stemless, small subshrubs, or cushion plants; indumentum of medifixed hairs. Leaves alternate,
pinnatifid or 3-lobed. Synflorescences cymose, dense or lax, forming panicles, clusters, or heads. Capitula few to many, homogamous,
discoid. Involucre campanulate or conical; phyllaries in 3 or 4 rows, herbaceous or rigidly herbaceous, scarious margins white or
brown. Receptacle flat to convex, epaleate. Florets all bisexual; corolla tubular, apex 5-lobed. Anther bases obtuse, apical appendage
ovate-lanceolate. Style branches linear, apex truncate. Achenes subterete, 4–7-ribbed, with a ± distinct apical rim. Corona absent.
Nineteen species: C Asia, China, Himalaya, Mongolia; 11 species (six endemic) in China.
Hippolytia dolichophylla (Kitamura) K. Bremer & Humphries (Bull. Nat. Hist. Mus. London, Bot. 23: 106. 1993; Chrysanthemum dolichophyllum
Kitamura, Acta Phytotax. Geobot. 23: 73. 1968, based on Tanacetum longifolium Wallich ex Candolle, Prodr. 6: 130. 1838, not Thunberg
(1800); C. longifolium K. H. Rechinger (1955), not Sessé & Mociño (1894); T. dolichophyllum (Kitamura) Kitamura; T. himachalense Aswal &
Mehrotra, nom. illeg. superfl.) was recorded from Sichuan in Annot. Checkl. Fl. Pl. Nepal (69. 2000, as T. dolichophyllum), disjunct from the main
area of distribution in the Himalaya between Kashmir and Nepal (see http://www.efloras.org/florataxon.aspx?flora_id=110&taxon_id=242425863;
accessed 8 Apr 2011). No specimen has been seen by the present authors. This species would key out as H. herderi, but it differs from that species and
all others in Hippolytia by having leaves 12–25 cm.