9. Tribe ANTHEMIDEAE

764
ANTHEMIDEAE
1a. Ray florets absent: capitula disciform or discoid.
2a. Florets orange-red; plants 7–25 cm tall ............................................................................................................. 10. T. tatsienense
2b. Florets yellow; plants 30–150 cm tall.
3a. Synflorescence with (5–)10–70(–100) capitula; stem leaves elliptic or elliptic-ovate, to 20 cm, ultimate
segments ovate, obliquely triangular, or narrowly elliptic; capitula disciform .................................................. 1. T. vulgare
3b. Synflorescence with 3–10(–12) capitula; stem leaves linear-oblong or oblong, 1–9 cm, ultimate segments
linear or oblong; capitula discoid.
4a. Leaves less than 3 cm, with inconspicuous or short petioles, ultimate segments oblong, apex
obtuse ................................................................................................................................................. 18. T. artemisioides
4b. Leaves more than 4 cm, with long petioles, ultimate segments linear, acute .......................................... 19. T. falconeri
1b. Ray florets conspicuous: capitula radiate.
5a. Ray florets yellow.
6a. Capitula 1 per stem; indumentum with curved simple hairs only ................................................................. 11. T. atkinsonii
6b. Capitula 3–16 per stem; indumentum with a mixture of simple hairs and T-shaped hairs.
7a. Basal leaves inconspicuously tripinnatisect, petiole to ca. 2.5 cm ....................................................... 6. T. scopulorum
7b. Basal leaves 2-pinnatisect, petiole 3–8 cm.
8a. Capitula 3–7, densely corymbose or compound-corymbose; peduncle becoming thick at apex,
0.5–1.5 cm ........................................................................................................................................... 3. T. crassipes
8b. Capitula 5–16, laxly irregularly corymbose; peduncle 1–10 cm.
9a. Leaves narrowly elliptic or narrowly linear-elliptic; involucre 7–17 mm in diam. .............. 4. T. barclayanum
9b. Leaves linear or broadly linear; involucre 4–7 mm in diam. ...................................................... 5. T. santolina
5b. Ray florets white, pink, red, orange-red, or purplish.
10a. Ray florets red, orange-red, or purplish.
11a. Plants 25–50 cm tall; basal leaves 2.5–4 cm wide; outer phyllaries ca. 3.5 mm .............................. 8. T. coccineum
11b. Plants 7–35 cm tall; basal leaves 1–2 cm wide; outer phyllaries 5–6 mm.
12a. Ray florets orange-red; corona ca. 0.1 mm, divided to base ................................................. 10. T. tatsienense
12b. Ray florets red or purple-red; corona 0.3–0.5 mm, divided for half length ........................ 16. T. richterioides
10b. Ray florets white or pink.
13a. Small subshrubs; ray florets pink ......................................................................................................... 9. T. petraeum
13b. Perennial herbs; ray florets white (sometimes pink in T. kaschgarianum).
14a. Stems and leaves with T-shaped or forked hairs, sometimes mixed with simple hairs.
15a. Basal leaves with petiole 6–9 cm, blade ca. 10 × 2.5 cm; stems and leaves green
or gray-green; indumentum spreading, hairs a mixture of simple and T-shaped
hairs; involucres campanulate ...................................................................................... 2. T. tanacetoides
15b. Basal leaves with petiole 10–20 cm, blade 1.5–4 × 1–2 cm; stems and leaves
silvery gray; indumentum appressed, all hairs T-shaped or forked; involucres
shallowly cup-shaped ............................................................................................... 7. T. cinerariifolium
14b. Stems and leaves with simple hairs only, sometimes these glandular.
16a. Involucres straw-colored; scarious margins of phyllaries narrowly white;
leaves petiolate (cultivated ornamentals).
17a. Plants green or deep green; ray floret lamina 7–10 mm;
achenes ca. 3 mm ........................................................... T. partheniifolium (see comment above)
17b. Plants yellow-green; ray floret lamina 3–6 mm; achenes
ca. 1.2 mm ............................................................................. T. parthenium (see comment above)
16b. Involucres not straw-colored; scarious margins of phyllaries
dark brown or brown; stem leaves sessile or shortly petiolate.
18a. Leaf surfaces gray-white; receptacle pilose with easily shed hairs ............. 17. T. kaschgarianum
18b. Leaf surfaces green; receptacle persistently pilose.
19a. Capitula 2–7, very rarely solitary.
20a. Indumentum with a mixture of glandular and eglandular hairs;
ultimate segments of leaves filiform ............................................. 14. T. abrotanoides
20b. Indumentum with glandular hairs only; ultimate segments of
leaves linear to obliquely triangular ................................................. 12. T. alatavicum
19b. Capitula solitary, very rarely 2 or 3.
21a. Basal leaves 1–2 cm wide; achene corona 1–1.2 mm, divided
to base .................................................................................................. 15. T. pulchrum
21b. Basal leaves 2–5 cm wide; achene corona 0.4–0.7 mm, divided
for half length ................................................................................. 13. T. krylovianum