FAO Species Identification Guide for Fishery Purposes Western

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Onychoteuthidae 785 Key to the genera of Onychoteuthidae occurring in the area 1a. Skin smooth; posterior end of gladius a short chitinous needle (Fig. 1a); light organs present: 1 elongated light organ on ventral side of eyeball, 2 light organs on intestine and ink sac (anterior one small, posterior one large) (Fig. 2a) . . . . . . . . . . . . . Onychoteuthis 1b. Skin smooth or rough; posterior end of gladius a long opaque cartilaginous rostrum (Fig. 1b); no light organs present (Fig. 2b) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . → 2 a) Onychoteuthis short needle Fig. 1 gladius long rostrum b) Ancistroteuthis, Moroteuthis nuchal folds or pleats intestine 2a. Six to 10 well-developed nuchal folds; skin smooth (Fig. 3) . . . . . . . . . . . . . Ancistroteuthis 2b. No nuchal folds (Fig. 4a); skin thick, rough, or tuberculate (Fig. 4b) . . . . . . . . . . . Moroteuthis dorsal view Fig. 3 Ancistroteuthis ink sac light organs a) head and anterior b) light organs on part of mantle intestinal tract (dorsal view) (ventral view) c) dorsal view Fig. 2 Onychoteuthis nuchal folds absent a) head and anterior part of mantle b) dorsal view (dorsal view) (illustration: K. Hollis/ABRS) Fig. 4 Moroteuthis
786 Cephalopods List of species occurring in the area Ancistroteuthis lichtensteini (Orbigny, 1839) Moroteuthis lönnbergi Ishikawa and Wakiya, 1914 Onychoteuthis “banksii” Leach, 1817 - species complex Onychoteuthis compacta (Berry, 1913) Onychoteuthis meridiopacifica Rancurel and Okutani, 1990 References Kubodera, T., U. Piatkowski, T. Okutani, and M.R. Clareke. 1996. Taxonomy and Zoogeography of the Family Onychoteuthidae (Cephalopoda: Oegopsida). Smithson. Contrib. Zool. Rancurel, P. 1976. Note sur les Cephalopodes des contenus stomacaux de Thunnus albacares (Bonnaterre) dans le Sud-ouest Pacifique. Cah. ORSTOM (Ser. Océanog.), 14:71-80. Tsuchiya, K and T. Okutani. 1991. Growth stages of Moroteuthis robusta (Verrill, 1881) with a re-evaluation of the genus. Bull. Mar. Sci., 49:137-147.
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click for previous page iv Table of
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vi Page Brachaeluridae . . . . . .
Page 5 and 6:
688 Cephalopods Introduction and Ge
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690 Cephalopods Principal PRINCIPAL
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692 Cephalopods To examine the cont
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694 Cephalopods Chitin(ous) - a hor
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696 Cephalopods Ink sac - the struc
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698 Cephalopods Spermatophore groov
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700 Cephalopods 6a. Fins small and
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702 Cephalopods 15a. Surface of man
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704 Cephalopods 23a. Arms short, ty
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706 Cephalopods Order SEPIIDA - Cut
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708 Cephalopods List of Familes LIS
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710 Cephalopods Key to the species
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712 Cephalopods Sepiolidae SEPIOLID
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714 Cephalopods Eupryma morsei (Ver
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716 Cephalopods Rossia bipapillata
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718 Cephalopods Sepiolina nipponens
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720 Cephalopods Sepiadarium kochii
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722 Cephalopods Spirulidae SPIRULID
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724 Cephalopods anterior last locul
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726 Cephalopods it may be necessary
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728 Cephalopods 4a. Five to 6 wine-
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730 Cephalopods 11a. Tentacular clu
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732 Cephalopods 23a. Hectocotylus w
Page 51 and 52: 734 Cephalopods List of species occ
Page 53 and 54: 736 Cephalopods Sepiidae Sepia acul
Page 55 and 56: click for previous page 738 Cephalo
Page 57 and 58: 740 Cephalopods Sepia brevimana Ste
Page 59 and 60: 742 Cephalopods Sepia esculenta Hoy
Page 61 and 62: 744 Cephalopods Sepia latimanus Quo
Page 63 and 64: 746 Cephalopods Sepia opipara (Ired
Page 65 and 66: 748 Cephalopods Sepia pharaonis Ehr
Page 69 and 70: 752 Cephalopods Sepia rozella (Ired
Page 71 and 72: 754 Cephalopods Sepia stellifera Ho
Page 73 and 74: 756 Cephalopods Sepiella inermis F
Page 75 and 76: 758 Cephalopods Sepia bartletti (Ir
Page 77 and 78: 760 Cephalopods Sepia mestus Gray,
Page 79 and 80: 762 Cephalopods Sepia vossi Khromov
Page 83 and 84: 766 Cephalopods Loligo reesi (Voss,
Page 85 and 86: 768 Cephalopods 6a. Gladius relativ
Page 87 and 88: 770 Cephalopods ventral view Fig. 1
Page 89 and 90: 772 Cephalopods Nipponololigo beka
Page 91 and 92: 774 Cephalopods “Photololigo chin
Page 93 and 94: 776 Cephalopods “Photololigo edul
Page 95 and 96: 778 Cephalopods “Sepioteuthis les
Page 97 and 98: 780 Cephalopods Loliolus affinis St
Page 99 and 100: 782 Cephalopods Similar families oc
Page 101: 784 Cephalopods Onychoteuthidae ONY
Page 107 and 108: 790 Cephalopods 4a. Mantle very sle
Page 109 and 110: 792 Cephalopods Nototodarus hawaiie
Page 111 and 112: 794 Cephalopods Sthenoteuthis ouala
Page 113 and 114: 796 Cephalopods Todaropsis eblanae
Page 115 and 116: 798 Cephalopods Chiroteuthidae CHIR
Page 119 and 120: 802 Cephalopods Ocythoidae (suborde
Page 121 and 122: 804 Cephalopods 5a. Distinctly elon
Page 123 and 124: 806 Cephalopods 14a. Small, elongat
Page 125 and 126: 808 Cephalopods 22a. Pale longitudi
Page 127 and 128: 810 Cephalopods Cistopus indicus (R
Page 129 and 130: 812 Cephalopods Octopus cyanea Gray
Page 131 and 132: 814 Cephalopods Octopus cf. luteus
Page 133 and 134: 816 Cephalopods Octopus nocturnus N
Page 135 and 136: 818 Cephalopods Octopus tetricus Go
Page 137 and 138: 820 Cephalopods Ameloctopus litoral
Page 139 and 140: 822 Cephalopods Octopus abaculus No
Page 141 and 142: 824 Cephalopods Octopus australis H
Page 143 and 144: 826 Cephalopods Octopus polyzenia G
Page 145 and 146: 828 Stomatopods Technical Terms and
Page 147 and 148: 830 Stomatopods propodus of HARPIOS
Page 149 and 150: 832 Stomatopods ODONTODACTYLIDAE Od
Page 151 and 152: 834 Stomatopods Odontodactylus scyl
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836 Stomatopods 2a. Antennal protop
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838 Stomatopods Harpiosquillidae HA
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840 Stomatopods 8a. Rostral plate l
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842 Stomatopods Squillidae SQUILLID
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844 Stomatopods 7a. Dorsal surface
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846 Stomatopods Cloridopsis scorpio
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848 Stomatopods Oratosquillina grav
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click for previous page SHRIMPS AND
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Technical Terms and Measurements 85
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General Remarks 855 The terms “sh
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List of Families 857 List of Famili
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Infraorder Penaeidae, Superfamily S
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Sergestidae 861 6a. Lower antennula
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Sergestidae 863 Acetes intermedius
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Sergestidae 865 Acetes vulgaris Han
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Superfamily Penaeoidea 867 Caridea:
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Aristeidae 869 Similar families occ
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Aristeidae 871 List of species occu
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Aristeidae 873 Aristeus virilis (Ba
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Solenoceridae 875 Solenoceridae SOL
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Solenoceridae 877 Stenopodidae: thi
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Solenoceridae 879 9a. Anterior end
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Solenoceridae 881 Haliporoides sibo
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Solenoceridae 883 Solenocera crassi
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Solenoceridae 885 Solenocera alfons
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Solenoceridae 887 Solenocera koelbe
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click for previous page Penaeidae 8
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Penaeidae 891 Sergestidae: size sma
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Penaeidae 893 9a. Carapace without
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Penaeidae 895 2a. Rostrum armed wit
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Penaeidae 897 17a. In males, basial
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Penaeidae 899 5a. Epigastric spine
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Penaeidae 901 2a. In males, petasma
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Penaeidae 903 9a. Adrostral crest e
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Penaeidae 905 Metapenaeopsis mannar
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Penaeidae 909 Metapenaeus anchistus
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Penaeidae 911 Metapenaeus intermedi
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Penaeidae 913 Parapenaeopsis hardwi
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Penaeidae 915 Penaeus esculentus Ha
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Penaeidae 917 Penaeus japonicus Bat
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Penaeidae 919 Penaeus longistylus K
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Penaeidae 921 Penaeus merguiensis D
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Penaeidae 923 Penaeus penicillatus
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Penaeidae 925 Penaeus semisulcatus
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Penaeidae 927 Trachypenaeus curviro
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Penaeidae 929 Atypopenaeus formosus
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Penaeidae 931 Metapenaeopsis novaeg
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Penaeidae 933 Metapenaeopsis wellsi
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Penaeidae 935 Metapenaeus conjunctu
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Penaeidae 937 Metapenaeus eboracens
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Penaeidae 939 Metapenaeus lysianass
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Penaeidae 943 Parapenaeopsis gracil
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Penaeidae 945 Parapenaeopsis tenell
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Penaeidae 947 Parapenaeus longipes
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Penaeidae 949 Trachypenaeus gonospi
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Penaeidae 951 Trachypenaeus villalu
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Sicyoniidae 953 Solenoceridae: body
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Stenopodidae 955 Stenopodidae Infra
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Infraorder Caridea 957 Infraorder C
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Infraorder Caridea 959 Superfamily
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Atyidae/Hippolytidae 961 Caridina w
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Hymenoceridae 963 Hymenoceridae HYM
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Palaemonidae 965 Exopalaemon stylif
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click for previous page Palaemonida
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Palaemonidae/Pandalidae 969 Palaemo
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Rhynchocinetidae 971 Rhynchocinetid
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974 Lobsters Technical Terms and Me
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976 Lobsters General Remarks GENERA
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978 Lobsters ENOPLOMETOPIDAE Page 9
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980 Lobsters List of FamiliesLIST a
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click for previous page 982 Lobster
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984 Lobsters Palinuridae: carapace
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986 Lobsters 6a. Abdomen smooth, wi
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988 Lobsters Acanthacaris tenuimana
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990 Lobsters Metanephrops thomsoni
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992 Lobsters Metanephrops andamanic
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994 Lobsters Metanephrops velutinus
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996 Lobsters Similar families occur
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998 Lobsters 3a. Carapace with 5 me
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1000 Lobsters Enoplometopus holthui
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1002 Lobsters Enoplometopidae: body
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1004 Lobsters Palibythus magnificus
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1006 Lobsters gonopore female leg 1
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1008 Lobsters Synaxidae: body very
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1010 Lobsters Key to the species of
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1012 Lobsters Key to the species of
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1014 Lobsters 8a. Antennular plate
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1016 Lobsters Panulirus homarus (Li
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1018 Lobsters Panulirus ornatus (Fa
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1020 Lobsters Panulirus polyphagus
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1022 Lobsters Justitia chani Poupin
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1024 Lobsters Linuparus sordidus Br
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1026 Lobsters Panulirus pascuensis
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click for previous page 1028 Lobste
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1030 Lobsters 5a. Medium size (adul
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1032 Lobsters 2a. Articulated parts
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1034 Lobsters Ibacus ciliatus (Von
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1036 Lobsters Ibacus pubescens Holt
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click for previous page 1038 Lobste
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1040 Lobsters Thenus orientalis (Lu
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1042 Lobsters Parribacus holthuisi
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click for previous page CRABS by P.
Page 361 and 362:
Technical Terms and Measurements 10
Page 363 and 364:
General Remarks 1049 Poisonous Crab
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General Remarks 1051 Adult male and
Page 367 and 368:
General Remarks 1053 longitudinally
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Imported Crabs of Commercial Import
Page 371 and 372:
Guide to Families 1057 CALAPPIDAE P
Page 373 and 374:
Guide to Families 1059 PORTUNIDAE P
Page 375 and 376:
Guide to Families/Key to Brachyura
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Key to Brachyura 1063 Fig. 9 Cyclod
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Key to Brachyura 1065 13a. Opening
Page 381 and 382:
Key to Brachyura 1067 18a. Antennae
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Key to Brachyura 1069 Fig. 46 Atele
Page 385 and 386:
Key to Brachyura 1071 30a. Last pai
Page 387 and 388:
Key to Brachyura 1073 strongly recu
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Key to Brachyura 1075 42a. Longitud
Page 391 and 392:
Key to Anomura 1077 48a. Male first
Page 393 and 394:
Key to Anomura 1079 a) Fig. 103 Coe
Page 395 and 396:
List of Families 1081 * CHEIRAGONID
Page 397 and 398:
click for previous page Infraorder
Page 399 and 400:
Dromiidae 1085 Dromiidae DROMIIDAE
Page 401 and 402:
Dromiidae 1087 Dromia dormia (Linna
Page 403 and 404:
Raninidae 1089 Raninidae RANINIDAE
Page 405 and 406:
Calappidae 1091 Calappidae CALAPPID
Page 407 and 408:
Calappidae 1093 3a. Entire posterio
Page 409 and 410:
Calappidae 1095 Matuta victor (Fabr
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Calappidae 1097 Calappa hepatica (L
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Xanthidae 1099 Key to species of in
Page 415 and 416:
Xanthidae 1101 Etisus splendidus Ra
Page 417 and 418:
click for previous page Eriphiidae
Page 419 and 420:
Eriphiidae 1105 6a. Dorsal surface
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Eriphiidae 1107 Menippe rumphii (Fa
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Eriphiidae 1109 Myomenippe fornasin
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Carpiliidae 1111 Carpilius convexus
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Pilumnidae 1113 A single species of
Page 429 and 430:
click for previous page Portunidae
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Portunidae 1117 6a. Posterior borde
Page 433 and 434:
Portunidae 1119 16a. Carapace with
Page 435 and 436:
Portunidae 1121 Charybdis japonica
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Portunidae 1123 Podophthalmus vigil
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Portunidae 1125 Portunus sanguinole
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Portunidae 1127 The differences in
Page 443 and 444:
Portunidae 1129 Thalamita crenata (
Page 445 and 446:
Portunidae 1131 Charybdis annulata
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Geryonidae 1133 3a. Anterolateral m
Page 449 and 450:
Geryonidae 1135 Chaceon australis M
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Majidae 1137 Schizophrys aspera (H.
Page 453 and 454:
Grapsidae 1139 2a. Carapace squaris
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Grapsidae 1141 Key to food species
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Grapsidae 1143 Episesarma versicolo
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Grapsidae 1145 Episesarma chengtong
Page 461 and 462:
click for previous page Gecarcinida
Page 463 and 464:
Gecarcinidae 1149 Cardisoma carnife
Page 465 and 466:
Gecarcinidae 1151 Cardisoma hirtipe
Page 467 and 468:
Ocypodidae 1153 Ocypode ceratophtha
Page 469 and 470:
Infraorder Anomura - Coenobitidae 1
Page 471 and 472:
1158 Holothurians General Remarks G
Page 473 and 474:
1160 Holothurians dentritic peltate
Page 475 and 476:
1162 Holothurians oral tentacles ge
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1164 Holothurians Interradii (or in
Page 479 and 480:
1166 Holothurians 11a. Tables with
Page 481 and 482:
1168 Holothurians Actinopyga maurit
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1170 Holothurians Actinopyga palaue
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1172 Holothurians Bohadschia argus
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1174 Holothurians Bohadschia vitien
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1176 Holothurians Holothuria (Halod
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1178 Holothurians Holothuria (Merte
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1180 Holothurians Holothuria (Metri
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1182 Holothurians Holothuria (Micro
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1184 Holothurians Pearsonothuria gr
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1186 Holothurians Stichopus chloron
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1188 Holothurians Stichopus variega
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1190 Holothurians Thelenota anax Cl
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1192 Hagfishes MYXINIDAE Hagfishes
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1194 Sharks Technical Terms and Mea
Page 509 and 510:
1196 Sharks General Remarks GENERAL
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1198 Sharks Key to Families KEY TO
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1200 Sharks 12a. No lobe and groove
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1202 Sharks 21a. First dorsal fin l
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1204 Sharks PRISTIOPHORIDAE: Sawsha
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1206 Sharks PSEUDOTRIAKIDAE: False
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click for previous page 1208 Sharks
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1210 Sharks Heptranchias perlo (Bon
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1212 Sharks Key to the species of E
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1214 Sharks Squatinidae: trunk much
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1216 Sharks 11a. Diagonal distance
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1218 Sharks 20a. Upper teeth relati
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1220 Sharks 29a. Denticles on sides
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1222 Sharks Etmopterus molleri Whi
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1224 Sharks Centrophorus niaukang T
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1226 Sharks Etmopterus brachyurus S
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1228 Sharks Euprotomicrus bispinatu
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1230 Sharks Squalus megalops (Macle
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1232 Sharks Squalus sp. F [Last and
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1234 Sharks Key to the species of P
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1236 Sharks Rays (Batoidea): pector
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1238 Sharks HETERODONTIDAE Bullhead
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1240 Sharks Heterodontus galeatus (
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1242 Sharks List of species occurri
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1244 Sharks Key to the species of B
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1246 Sharks Similar families occurr
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1248 Sharks Orectolobus ornatus (de
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1250 Sharks Ginglymostomatidae: pre
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1252 Sharks 7a. Black spot behind g
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1254 Sharks Chiloscyllium hasselti
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1256 Sharks Chiloscyllium plagiosum
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1258 Sharks Hemiscyllium freycineti
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1260 Sharks GINGLYMOSTOMATIDAE Nurs
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1262 Sharks A single species in thi
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1264 Sharks Odontaspididae ODONTASP
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1266 Sharks Carcharias taurus Rafin
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1270 Sharks Key to the species of A
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1272 Sharks Alopias superciliosus (
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1274 Sharks LAMNIDAE Mackerel shark
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1276 Sharks Carcharodon carcharias
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1278 Sharks Isurus paucus Guitart M
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1280 Sharks Key to the species of S
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1282 Sharks 13a. Pectoral fin broad
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1284 Sharks 25a. Labial furrows ver
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1286 Sharks Apristurus herklotsi (F
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1288 Sharks Atelomycterus macleayi
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1290 Sharks Galeus gracilis Compagn
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1292 Sharks Scyliorhinus garmani (F
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1294 Sharks Triakidae: no gill rake
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1298 Sharks Proscylliidae: gill rak
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1300 Sharks 9a. Snout bluntly round
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1302 Sharks Galeorhinus galeus (Lin
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1304 Sharks Mustelus antarcticus G
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1306 Sharks Similar families occurr
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1308 Sharks Chaenogaleus macrostoma
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1310 Sharks Hemipristis elongatus (
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click for previous page 1312 Sharks
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1314 Sharks Odontaspididae: fifth g
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1316 Sharks 6a. Second dorsal-fin o
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1318 Sharks 12a. Head very flat and
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1320 Sharks 22a. Second dorsal fin
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1322 Sharks 30a. Usually 11 lower a
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1324 Sharks List of species occurri
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1326 Sharks Carcharhinus altimus (S
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1328 Sharks Carcharhinus amblyrhync
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1330 Sharks Carcharhinus borneensis
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1332 Sharks Carcharhinus brevipinna
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1334 Sharks Carcharhinus dussumieri
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1336 Sharks Carcharhinus fitzroyens
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1338 Sharks Carcharhinus hemiodon (
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1340 Sharks Carcharhinus limbatus (
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1342 Sharks Carcharhinus macloti (M
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1344 Sharks Carcharhinus obscurus (
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1346 Sharks Carcharhinus sealei (Pi
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1348 Sharks Carcharhinus tilstoni (
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1350 Sharks Lamiopsis temmincki (M
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1352 Sharks Negaprion acutidens (R
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1354 Sharks Rhizoprionodon acutus (
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1356 Sharks Rhizoprionodon taylori
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1358 Sharks Triaenodon obesus (Rüp
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1360 Sharks Glyphis sp. B En - Born
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1362 Sharks 2a. Anterior margin of
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1364 Sharks Sphyrna lewini (Griffit
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1366 Sharks Sphyrna zygaena (Linnae
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1368 The Living Marine Resources of
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FAO SPECIES IDENTIFICATION GUIDE FO
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Carpenter, K.E.; Niem, V.H. (eds) F
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FAO Species Identification Guide for Fishery Purposes Western

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