FAO Species Identification Guide for Fishery Purposes Western

732 Cephalopods
23a. Hectocotylus with 3 rows of normal suckers proximally; cuttlebone anterior striae
inverted U-shape (Fig. 29a). . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Sepia aculeata
23b. Hectocotylus with 5 to 6 rows of normal suckers proximally; cuttlebone anterior striae
inverted V-shape (Fig. 29b) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Sepia esculenta
24a. Tentacular club with 6 or less suckers in transverse rows . . . . . . . . . . . . . . . . . . . . . .→25 [Ghofar (1989) reports Sepia bandensis as having 7 to 9 suckers in transverse rows on the tentacular club. The
number of suckers in transverse rows cited in the original description of Adam (1939), and by subsequent workers
e.g. Okutani (1995), and followed here is 5]
24b. Tentacular club with 8 suckers in transverse rows . . . . . . . . . . . . . . . . . . . . . . . . . → 31
25a. Tentacular club with 3 or 4 suckers in transverse rows . . . . . . . . . . . . . . . . Sepia opipara
25b. Tentacular club with 5 or 6 suckers in transverse rows . . . . . . . . . . . . . . . . . . . . . .→ 26
26a. Cuttlebone without spine, or spine reduced to a tiny tubercle . . . . . . . . . . . . . . . . . .→ 27
26b. Cuttlebone with spine . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .→ 28
27a. Maximum size 500 mm mantle length; anterior edge of inner cone with a V-shaped
calcareous callus, or rough thickening posteriorly (Fig. 30a) . . . . . . . . . . . . . . Sepia apama
27b. Maximum size 50 mm mantle length; cuttlebone inner cone without calcareous callus
posteriorly (Fig. 30b) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Sepia bandensis
anterior striae
inverted
U-shape
a) Sepia aculeata b) Sepia esculenta
Fig. 29 cuttlebone (ventral view)
anterior
striae
inverted
V-shape
28a. Male arms I to IV with tetraserial
suckers cuttlebone inner cone limbs
of uniform width; cuttlebone sulcus
distinct, flanked by rounded ribs
(Fig. 31a, b) . . . . . . . . . . . . . . . . .→ 29
28b. Male arms I to III with tetraserial
suckers proximally, biserial suckers
distally; cuttlebone inner cone limbs
broaden posteriorly; cuttlebone sulcus
indistinct, not flanked by
rounded ribs (Fig. 32a, b) . . . . . . . . . .→ 30
calcareous
callus
a) Sepia apama
b) Sepia bandensis
Fig. 30 cuttlebone (ventral view)
(illustration: K. Hollis/ABRS)
anterior
striae
inverted
U-shape
sulcus
shallow
but distinct
a) Sepia latimanus b) Sepia plangon
Fig. 31 cuttlebone (ventral view)
anterior
striae
inverted
V-shape
sulcus
deep