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FAO Species Identification Guide for Fishery Purposes Western

FAO Species Identification Guide for Fishery Purposes Western

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1312 Sharks<br />

Carcharhinidae CARCHARHINIDAE<br />

Requiem sharks<br />

(also, ground sharks, blue sharks, sharpnose sharks)<br />

by L.J.V. Compagno and V.H. Niem<br />

Diagnostic characters: Small to large sharks. Trunk and precaudal tail cylindrical, not depressed and<br />

without lateral ridges: precaudal tail much shorter than trunk. Head not expanded laterally, conical to<br />

moderately depressed; 5 small- to medium-sized gill slits present, the last 1 to 3 over or behind pectoral-fin<br />

origins, their upper ends not expanded onto dorsal surface of head; no gill sieves and usually no gill rakers<br />

on internal gill slits (short dermal gill rakers present in Prionace); spiracles usually absent (but always<br />

present in Galeocerdo); nostrils well separated from mouth, without barbels, nasoral grooves, or circumnarial<br />

grooves; eyes on sides of head, with a well-developed nictitating lower eyelid; snout short to<br />

moderately long, conical and slightly pointed to depressed and broadly rounded, never greatly flattened<br />

and blade-like and without lateral teeth and barbels; mouth usually large, arched and elongated, and<br />

extending well behind eyes; labial furrows usually present on both jaws but generally greatly reduced,<br />

confined to mouth corners, and barely visible when mouth is closed (but Galeocerdo and Rhizoprionodon<br />

species have well-developed labial furrows); upper labial furrows usually not reaching front of mouth<br />

(except in Galeocerdo); teeth small to large, blade-like, with a single cusp and cusplets variably<br />

developed; anterior teeth in upper jaw smaller than lateral teeth and not separated from them by<br />

smaller intermediate teeth on each side. Two dorsal fins, without spines, the first dorsal fin moderately<br />

large, high and angular or subangular, much shorter than the caudal fin, its base located over the interspace<br />

between pectoral and pelvic-fin bases and entirely anterior to origins of pelvic fins (free rear tip of dorsal<br />

fin may reach or extend posterior to pelvic-fin origins in Scoliodon, Negaprion, Rhizoprionodon, and<br />

Triaenodon); second dorsal fin varying from less than 1/5 the height of the first dorsal fin to about as high<br />

as the first (Lamiopsis and Negaprion); anal fin present, moderately large, with its origin varying from<br />

somewhat anterior to the second dorsal-fin origin to under the first half of second dorsal-fin base; caudal<br />

fin strongly asymmetrical, much less than 1/2 of total length, with a rippled or undulated dorsal margin,<br />

a well-marked subterminal notch, and a short but well-defined lower lobe; vertebral axis of caudal fin<br />

raised above body axis. Caudal peduncle not strongly depressed dorsoventrally or widely expanded<br />

laterally with weak longitudinal keels (Prionace, Galeocerdo) or none; precaudal pits present and well<br />

developed. Intestinal valve of scroll type. Colour: brown, grey, yellowish or bluish above, white to cream<br />

or yellowish below, some species with prominent dark or light markings on fins; body usually without a<br />

prominent colour pattern (except <strong>for</strong> Galeocerdo).<br />

posterior margin<br />

apex of 1 st dorsal fin<br />

inner margin<br />

nictitating membrane<br />

nostril<br />

internasal<br />

width<br />

anterior<br />

nasal flap<br />

labial folds<br />

length of snout<br />

width of<br />

mouth<br />

inner margin<br />

of pectoral fin<br />

position of interdorsal<br />

ridge when present<br />

rolled<br />

unrolled<br />

intestinal valve of scroll type<br />

second dorsal fin<br />

precaudal pit<br />

free tip of<br />

posterior lobe<br />

subterminal<br />

notch<br />

dorsal view of caudal fin<br />

example of upper and lower teeth<br />

(blade-like, with a single cusp, often serrated)

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