FAO Species Identification Guide for Fishery Purposes Western

Grapsidae 1141
Key to food species of Varuninae occurring in the area
1a. Distal process of male first gonopod
with 1 process distinctly longer than
the other (Fig. 12a); occurs mainly in
continental shelf waters . . . . . . . . Varuna yui
1b. Distal process of male first gonopod
with both processes subequal in size
(Fig. 12b); occurs mainly in areas
with more oceanic waters . . . . Varuna litterata
List of species of interest to fisheries occurring in the area
The symbol is given when species accounts are included.
Subfamily GRAPSINAE
Grapsus albolineatus Lamarck, 1818
Grapsus tenuicrustatus (Herbst, 1783)
Subfamily PLAGUSIINAE
Plagusia tuberculata Lamarck, 1818
Subfamily SESARMINAE
Episesarma mederi (A. Milne Edwards, 1854)
Episesarma chengtongense (Serène and Soh, 1967)
Episesarma palawanense (Rathbun, 1914)
Episesarma singaporense (Tweedie, 1936)
Episesarma versicolor (Tweedie, 1940)
Subfamily VARUNINAE
Varuna litterata (Fabricius, 1798)
Varuna yui Hwang and Takeda, 1984
a) Episesarma
chengtongense
broad
truncate
tip
b) Episesarma
palawanense
broad
rounded
tip
c) Episesarma
versicolor
relatively
narrow
truncate tip
Fig. 11 male first gonopod
d) Episesarma
singaporense
relatively
broad,
bent tip
unequal processes
e) Episesarma
mederi
narrow,
bent
tip
less unequal processe
a) Varuna yui b) Varuna litterata
Fig. 12 male first gonopod
References
Alcock, A. 1900. Materials for a carcinological fauna of India. No. 6. The Brachyura Catometopa or Grapsoidea. J. Asiat.
Soc. Bengal, 69, pt. 2(3):279-486.
Crosnier, A. 1965. Crustacés Décapodes. Grapsidae et Ocypodidae. Faune de Madagascar, 18:1-143.