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FAO Species Identification Guide for Fishery Purposes Western

FAO Species Identification Guide for Fishery Purposes Western

FAO Species Identification Guide for Fishery Purposes Western

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748 Cephalopods<br />

Sepia pharaonis Ehrenberg, 1831<br />

Frequent synonyms / misidentifications: Crumenasepia<br />

hulliana Iredale, 1926;C. ursulae Adam, 1939;Sepia rouxii<br />

Férussac and d’Orbigny, 1841; S. <strong>for</strong>mosana Berry, 1912;<br />

S. tigris Sasaki, 1929 / None.<br />

<strong>FAO</strong> names: En - Pharaoh cuttlefish; Fr - Seiche<br />

pharaon; Sp - Sepia faraónica.<br />

Diagnostic characters: Mantle with a<br />

row of elongated tubercles along<br />

silvery stripe at base of fins, or<br />

covered with numerous small<br />

papillae. Head tubercles absent. Left<br />

ventral arm (IV) of males hectocotylized:<br />

10 rows of normal suckers<br />

proximally, followed by 6 rows of<br />

reduced suckers, then suckers<br />

normal to arm tip; reduction marked,<br />

2 dorsal series, smaller than 2<br />

ventral serries, oral surface wide,<br />

fleshy, with transversely grooved<br />

ridges. Tentacular club long; sucker-<br />

bearing face flattened; with 8suckers<br />

in transverse rows, differing<br />

markedly in size (5 or 6 median<br />

suckers enlarged); dorsal and<br />

ventral view<br />

cuttlebone<br />

dorsal view<br />

ventral protective membranes not fused at base of club, joined to stalk at base of club,<br />

not separated from stalk by a membrane. Buccal membrane with few, minute suckers.<br />

Cuttlebone median rib present, distinct, wider anteriorly; lateral ribs indistinct; sulcus<br />

extending entire length, flanked by rounded ribs; anterior striae inverted U-shape;<br />

inner cone limbs thickened posteriorly, <strong>for</strong>m a shiny swelling; outer cone narrow<br />

anteriorly, broadens posteriorly; spine without keel. Colour: head with a vivid,<br />

transverse striped pattern; dorsal mantle pale brownish with white blotches or spots,<br />

or pale brown with transverse saddle mark (in females), or with vivid transverse striped<br />

pattern (especially in males); fins with pale reflective line along base (sometimes<br />

broken), line is blue in females; arms vivid with transverse stripes.<br />

Size: Maximum mantle length 420 mm; maximum weight 5 kg. In the Gulf of<br />

Carpentaria (Australia), the largest male and female collected in 1990 and 1991<br />

surveys were 192 mm and 173 mm mantle length, respectively. In the Gulf of Thailand,<br />

the maximum size of animals caught is <strong>for</strong> males 350 mm mantle length, and females<br />

300 mm mantle length.<br />

Habitat,biology,and fisheries: A neritic demersal species, at depths from 10 to 110 m.<br />

In the Gulf of Thailand and the Andaman Seas, found from the coastal shallows to a<br />

tentacular<br />

club<br />

(after Lu, in press)<br />

depth of 100 m, with most caught between 10 and 40 m. In the Gulf of Thailand, spawning occurs all year<br />

round, with peak months of January-February and July-September. Males mature at 13.7 mm mantle<br />

length, females at 142 mm mantle length. Supports industrial or artisanal fisheries throughout its range.<br />

With Sepia esculenta, the most abundant cuttlefish species caught in the Philippines. Also important to the<br />

commercial squid fishery of Thailand. Contributes about 90% of the cuttlefish catch off Australia by Chinese<br />

pair trawlers. This amounted to<br />

some 1 000 t in 1979. Taken by<br />

domestic fisheries as bycatch of<br />

prawn and mixed species trawl<br />

fisheries. The flesh is thick, tender<br />

and excellent <strong>for</strong> human consumption.<br />

Distribution: South and East<br />

China seas, Indonesia, Malaysia,<br />

Philippines, northern Australia, Gulf<br />

of Suez, Zanzibar, Madagascar to<br />

the Arabian Sea.<br />

10 mm

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