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Dissertation - HQ

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Choice of settlement habitat 45<br />

larvae arriving off a reef during the daytime when they only settle at<br />

night. Would these larvae simply continue on, past suitable habitat,<br />

or would they sense the presence of the habitat and behave in a way<br />

that keeps them in the vicinity until nightfall? There is little direct<br />

information on this sort of behaviour, although circumstantial evidence<br />

indicates that larvae do accumulate in the vicinity of settlement habitat<br />

to wait for the appropriate time 142 . This circumstantial evidence does<br />

not, however, help to decide over what periods of time such accumulation<br />

might take place. Information on the swimming, orientation or<br />

sensory abilities of the species of interest can be used to eliminate from<br />

consideration accumulation that is beyond the capabilities of the larvae.<br />

When habitat quality is known to influence settlement probability, it<br />

should be represented in the model, either explicitly when the fineness<br />

of the grid permits, or through parameterisation in each grid cell (e.g.<br />

description of the percentage of each habitat type in the cell). Settlement<br />

probability should be deduced from this habitat map. Caution is advised<br />

regarding how this probability is computed. One could be tempted<br />

to use settlement patterns observed a posteriori to compute a spatial<br />

probability of settlement. While this would be likely to enhance the<br />

result of the model when compared to observations, it does not rely on<br />

a mechanistic understanding of the process, hence has no predictive<br />

value. Instead, settlement experiments should be done in all habitat<br />

types and probabilities should be derived from those.<br />

Spatial probability<br />

of settlement

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