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Certificate - Etheses - Saurashtra University

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Marchinton, 1980; Smith, 1986; Hellgren and Vaughan, 1989; Oli et al., 1997; Dobey<br />

et al., 2002 and Hersey et al., 2005).<br />

2.6 Feeding ecology<br />

Most bears were opportunistic omnivores, their diets comprised of fruits, other<br />

vegetative materials, and in lesser amounts, mammals, fishes and insects.<br />

Evolutionary, brown bears have developed several adaptations for herbivory,<br />

including expansion of molar chewing surfaces and longer claws for digging.<br />

Nevertheless, they have maintained an unspecialized digestive system capable of<br />

digesting protein with efficiency equal to that of obligate carnivores (Bunnell and<br />

Hamilton, 1983). Most commonly, brown bear feeding habits have been quantified by<br />

analysis of scat contents. However, because of the differential digestibility of foods,<br />

contents of fecal residue were rarely equivalent to amounts of foods ingested by bears.<br />

The resulting underestimation of highly digestible foods was found most pronounced<br />

for meat and fish diets (Hewitt and Robbins, 1996).<br />

The major food items of brown bear were grouped into variety of ways based on<br />

taxonomic group and method of acquisition (LeFranc et al., 1987). They included (1)<br />

vegetative matter readily available for grazing such as graminoids, horsetails and<br />

forbs; (2) roots, corns and bulbs acquired by digging; (3) fruits produced from shrubs;<br />

(4) insects harvested from nests or aggregation sites, including ants (Formicidae),<br />

wasps (Vespidae) and moths and beetles; (5) mammals and birds, acquired through<br />

predation or scavenging, including ungulates and rodents and (6) fish acquired<br />

through predation or scavenging, including salmon and trout.<br />

21

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