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Arkell.1956.Jurassic..

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34 THE BRITISH ISLES<br />

much more satisfactorily than in the past, for in the Jura the ammonite<br />

faunas of the Saxon and English basins meet and are combined in one<br />

succession. A gap in the English sequence exists between the Scissum<br />

and Murchisonae Zones, but it has turned out to be not so great as was<br />

supposed from the Hanover succession, where six subzones were intercalated<br />

between opalinum and murchisonae by Hoffmann (1913) (see below,<br />

p. 144), and it is probably bridged to some extent by the Ancolioceras<br />

beds of Buckman, which represent some of the lower parts of the<br />

Murchisonae Zone. There are visible signs of pre-Murchisonae Zone<br />

erosion in Yorkshire.<br />

ZONES SUBZONES<br />

COTSWOLD<br />

FORMATIONS<br />

Graphoceras concavum Graphoceras concavum (Sow.) -| Tilestone<br />

Snowshill Clay<br />

Harford Sands<br />

Ludwigia murchisonae •<br />

Brasilia bradfordensis Buckman -J<br />

Ludwigia murchisonae (Sow.) j<br />

'Ancolioceras spp.'<br />

Upper Freestone<br />

Oolite Marl<br />

Lower Freestone<br />

Pea Grit<br />

? Lower Limestones<br />

Tmetoceras scissum Tmetoceras scissum (Benecke) Scissum Beds<br />

Leioceras opalinum Leioceras opalinum (Reinecke) (absent)<br />

The Scissum Zone is upheld as a separate zone because it is distinct<br />

in the Jura (where it is found to be contemporaneous with the German<br />

subzone of Costileioceras sinon Bayle sp.) and has been recognized in<br />

Canada and Argentina. It therefore has wide stratigraphical significance,<br />

despite the isolated rare occurrence of a minute Tmetoceras in the condensed<br />

Bradfordensis and Concavum Bed in Dorset (Bomford, 1948).<br />

Buckman and Richardson traced the Scissum Zone as a distinct formation<br />

over most of the outcrops of southern England. Its discrimination as a<br />

separate zone between those of Opalinum and Murchisonae is due to<br />

Buckman (1898), but Tmetoceras scissum had already been proposed as<br />

index species for a Mediterranean zone, supposed to be equivalent to<br />

these two northern zones, by Neumayr (1871, Der Penninische Klippenzug,<br />

p. 509).<br />

The Scissum and Opalinum Zones acquire great economic importance<br />

in the English Midlands, where they are developed as the Northampton<br />

Sand, with its valuable low-grade ironstone. The ammonites of this<br />

include Leioceras opalinum and Tmetoceras scissum, with Bredyia and<br />

Lytoceratids (Hollingworth & Taylor, 1946, 1951).<br />

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