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LOWER OXFORDIAN<br />

RUSSIA 497<br />

Cordatum Zone. Bed Ai, with Cardioceras ilovaiskyi Maire (Ilov.,<br />

pi. x, fig. 28) and other forms of the cordatum group (pi. x, fig. 31),<br />

Goliathiceras sp. (pi. x, fig. 27), Pachycardioceras rouilleri (p. 268) and<br />

Perisphinctids. Other ammonites of the Cordatum Zone were figured by<br />

Lahusen (1883): C. cordatum (Sow.) (pi. v, fig. 5), C. costicardia (Buck.)<br />

(pi. v, fig. 3), Aspidoceras aff. catena (Sow.) (pi. x, fig. 14), Peltoceratoides<br />

sp. (pi. x, fig. 12).<br />

Mariae Zone. Bed K contains Quenstedtoceras (Pavloviceras) pavlowi<br />

Douville, of which the type came from Ryazan, and allied species<br />

(Ilovaisky, pi. x, figs. 22-25; Lahusen, 1883, pi. iv, figs. 6-17). To the<br />

same zone belongs Cardioceras (Scarburgiceras) lahuseni Maire (Lahusen,<br />

1883, pi. v, fig. 1). At Miatchkovo near Moscow, this zone rests on<br />

Carboniferous Limestone, which is quarried on both banks of the Volga.<br />

UPPER CALLOVIAN<br />

The Lamberti Zone is strongly represented on the Upper Volga and in<br />

the Ryazan province by the Leachi Zone of Nikitin, with a range of<br />

Quenstedtoceratids comparable with those in the Hackness Rock and the<br />

Lamberti Limestone at Woodham pit in England. The clay of this zone<br />

similarly passes into a band of argillaceous limestone in places on the<br />

upper Volga; and at Novosselki in Ryazan province it oversteps on to<br />

Carboniferous Limestone. A notable link with England is the presence<br />

of conspicuous large Eboraciceras, to which belong Q. rybinskianum<br />

Nik. (1881, pi. i, fig. 8) and Q. problematicum Sinzow, 1899 (type, Nikitin,<br />

1881, pi. i, fig. 6), together with Lamberticeras (pi. i, fig. 1, and Lahusen,<br />

1883, pi. iv, figs. 1-3, and perhaps fig. 4, the type of Q. pseudolamberti<br />

Sinzow, 1899), Prorsiceras (Nikitin, 1881, pi. i, fig. 5), Pachyceras mologae<br />

Nikitin sp. (pi. i, fig. 12), and the ubiquitous Kosmoceras spinosum Sow.<br />

sp. (pi. iv, fig. 34) and K. proniae and its allies (Teisseyre, 1884). The<br />

last and K. transitionis Nik. and various Peltoceratids (Lahusen, 1883,<br />

pi. x, figs. 5-9) indicate that the Athleta Zone is represented. Various<br />

well-preserved Hecticoceratids come from the Upper and Middle Callovian<br />

(Lahusen, pi. xi).<br />

MIDDLE CALLOVIAN<br />

The Coronatum and Jason Zones are widespread and are represented<br />

by Lahusen's beds b, c. They yield Erymnoceras coronatum and E. doliforme<br />

Roman (Neumayr, 1876, pi. xxv, figs. 1-3; Lahusen, 1883, pi. vi, figs. 1-4)<br />

and numerous appropriate Kosmoceras, Hecticoceras, etc. (Lahusen,<br />

pis. vii-xi). Presumably from this level also comes Erymnoceras renardi<br />

Nikitin sp. (1882, pi. xi, fig. 24). At Sergatch the Coronatum Zone with<br />

Erymnoceras is developed as a band of oolite (Milachewitch, 1880).<br />

Classic works, chiefly based on ammonites from this substage in Ryazan<br />

province, were published by Neumayr (1876) and Teisseyre (1884).<br />

According to Zonov (1937, p. 35) Erymnoceras spp. occur in the lower part<br />

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