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ORCHIDACEAE - China

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ORCHIDOIDEAE 63lip ca. 1.6 cm, 3-partite; hypochile saccate and shortly spurred,ca. 2 mm, inside with 2 3- or 4-lobed calli; mesochile ca. 6 mm;epichile with 2 green spots at base, deeply bilobed; lobes ca. 10× 7 mm, 4- or 5-laciniate, each division 1.5–2.5 mm. Columnca. 2.5 mm; stelidia ca. 4 mm, slightly longer than rostellumarm. Fl. Mar.● Forests on limestone mountains; ca. 1200 m. S Yunnan.11. KUHLHASSELTIA J. J. Smith, Icon. Bogor. 4: 1, t. 301. 1910.旗 唇 兰 属 qi chun lan shuChen Xinqi ( 陈 心 启 Chen Sing-chi); Stephan W. Gale, Phillip J. CribbVexillabium F. Maekawa.Herbs, terrestrial. Rhizome elongate, creeping, noded, fleshy. Stem ascending, glabrous, leafy. Leaves ovate to subcircular, witha short petiole-like base sheathing stem. Inflorescence erect, pubescent; peduncle green, sometimes tinged with purplish red, withscattered sterile bracts; rachis 1- to many flowered; floral bracts lanceolate, ca. as long as pedicel and ovary, membranous, oftenpubescent and with ciliate margin. Flowers resupinate, small; ovary twisted, glabrous to pubescent. Sepals adaxially glabrous topubescent; lateral sepals connate along inner margin for up to half their length, forming a tube and enclosing hypochile. Petals membranous,upper margin adnate to dorsal sepal and forming a hood; lip longer than sepals, attached at base to column margin, 3-partite;hypochile a shallowly bilobed sac, sometimes septate, containing 2 oblong appendages; mesochile elongate, margin involute, subtubular,with a fleshy flange along exterior margin on each side; epichile dilated, flabellate, obcordate to transversely oblong, entireor bilobed. Column erect, stout, basally dilated; anther on ventral side of column, ovoid, 2-locular; pollinia 2, clavate, each longitudinallyparted, granular-farinaceous, sectile, with short caudicle, attached to a solitary viscidium; rostellum at column apex, erect,2-lobed; stigma lobes connate, at base of rostellum.About ten species: Indonesia, Malaysia, New Guinea, and the Philippines, north to <strong>China</strong>, Japan, and Korea; one species in <strong>China</strong>.Recent analyses of the Goodyerinae by Ormerod (Lindleyana 17: 189–238. 2002) and Pridgeon et al. (Gen. Orchid. 3: 63–153. 2003) have concludedthat Vexillabium and Kuhlhasseltia are congeneric.1. Kuhlhasseltia yakushimensis (Yamamoto) Ormerod, Lindleyana17: 209. 2002.旗 唇 兰 qi chun lanAnoectochilus yakushimensis Yamamoto, Bot. Mag. (Tokyo)38: 131. 1924; Cystopus humilis Fukuyama; Kuhlhasseltiaintegra (Fukuyama) T. C. Hsu & S. W. Chung; Pristiglottishumilis (Fukuyama) Fukuyama; P. integra Fukuyama; P. yakushimensis(Yamamoto) Masamune; Vexillabium humilum(Fukuyama) S. S. Ying; V. integrum (Fukuyama) S. S. Ying; V.yakushimense (Yamamoto) F. Maekawa.Plants 8–15 cm tall. Rhizome branching, pubescent. Stemascending, green. Leaves subrosulate at base of stem or spacedalong stem, green, tinged with reddish purple, ovate, 0.8–5 ×0.6–2.5 cm, fleshy, 3-veined, base rounded, margin sometimesminutely toothed, apex acute; petiole-like base 5–15 mm. Inflorescenceoften purplish red, 4–8 cm, white pubescent, with 1–3pinkish red sterile bracts; rachis pinkish, 1.5–4.5 cm, 2–7-flowered,sparsely pubescent; floral bracts purplish red, broadly lanceolate,5–6 mm, abaxially sparsely pubescent, margin ciliate,apex acuminate. Flowers small; ovary slightly curved, cylindric-fusiform,7–8 mm including pedicel, glabrous to pubescent.Sepals green at base, whitish pink toward apex; dorsalsepal shortly connate to lateral sepals at base, erect, oblongovate,concave, 3.5–4.5 mm, 1-veined, apex obtuse; lateralsepals oblong, falcate, 5–6 × ca. 2 mm, 1-veined, apex obtuse.Petals ovate, falcate, 4–5 × 1.5–2.5 mm, apex obtuse-apiculate;lip white, ca. 8 mm; hypochile saccate, containing 2 appendages;mesochile with 2 usually dentate flanges; epichile divergently2-lobed, 3–4 × 4.5–5 mm. Column 2–3 mm; anther cordate,ca. 1.4 mm, apex acuminate; pollinia obovoid, attached totriangular viscidium; rostellum erect, furcately 2-lobed, lobesunequal in size. Fl. Aug–Sep. 2n = 26.Forests, rocky crevices, along streams; 400–1600 m. Anhui,Hunan, Shaanxi, Sichuan, Taiwan, Zhejiang [Japan, Philippines].The S Taiwan (Lan Yu) endemic Pristiglottis integra has a pubescentovary but in other character states overlaps with Kuhlhasseltiayakushimensis.12. MYRMECHIS (Lindley) Blume, Coll. Orchid. 76. 1859.全 唇 兰 属 quan chun lan shuChen Xinqi ( 陈 心 启 Chen Sing-chi); Stephan W. Gale, Phillip J. CribbAnoectochilus sect. Myrmechis Lindley, Gen. Sp. Orchid. Pl. 500. 1840; Tubilabium J. J. Smith.Herbs, terrestrial or rarely epiphytic, small. Rhizome elongate, creeping or decumbent, several noded, fleshy; roots hairlike,arranged in irregular clumps or ridges along rhizome internodes. Stem ascending, terete, glabrous, with few to many scattered leaves.Leaves green, ovate or orbicular, small, usually less than 2 cm, slightly fleshy, base shortly petiolate and dilating into tubularamplexicaul sheath. Inflorescence abbreviate or occasionally elongate, glabrous to pubescent, with a few sheathing bracts proximallyand 1 to few flowers in a short terminal raceme. Flowers not opening fully, resupinate, small; ovary erect, twisted, glabrous to pubes-

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