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CAD. LAB. XEOL. LAXE 26 (2001) The evolution of metapodial bones 371 REFERENCES KUNST, G. K. (1992). Hoch- und spätglaziale Großsäugerreste aus dem Nixloch bei L o s e n s t e i n - Te r n b e rg (O.Ö.). Mitt. Komm. Quartärforsch., 8: 83-127, Wien, GUZVICA, G. & RADANOVIC–GUZVICA, B. (2000). Comparative-–evolutional Analyses of Cave Bear Metapodial Bones from North- Western Croatia. Geol. zbor., 1 5: 17-19, Ljubljana. MOTTL, M. (1947). Die Repolusthöhle, eine Protoaurignacienstation bei Peggau in der Steiermark. Verh. Geol. Bundesanst ., 1947: 200- 205, Wien. OSWALD, J. (1999). Geschlechtsdimorphismen an den Eckzähnen von Höhlenbären. Unpubl. Diplomarbeit, 51 S., Formal- und Naturwiss. Fakultät, Universität Wien. RABEDER, G. (1999). Die Evolution des Höhlenbärengebisses. Mitt. Komm. Quartärforsch., 11: 1-102, Wien. RABEDER, G & NAGEL, D. (2001). Phylogeny and new taxonomy of Alpine cave bears. This volume. THENIUS, E., (1956). Zur Kenntnis der fossilen Braunbären (Ursidae, Mammalia). Anz. Österr. Akad. Wiss., math.-naturw. Kl., Abt. I, 165: 153- 172, Wien. TORRES PEREZ-HIDALGO, T. de, (1988). Osos (Mammalia, Carnivora, Ursidae) del Pleistoceno de la Peninsula Ibérica. B o l e t í n Geológico y Minero, X C I X / I – V I : 1 - 3 1 6 , Instituto Geológico y Minero de España, Madrid.
Cadernos Lab. Xeolóxico de Laxe Coruña. 2001. Vol. 26, pp. 373-398 ISSN: 0213-4497 Pleistocene small cave bear (Ursus rossicus) from the South Siberia, Russia Un pequeño Oso de las Cavernas (Ursus rossicus) del Sur de Siberia, Rusia A B S T R A C T BARYSHNIKOV, G. 1 , FORONOVA, I. 2 The skull, mandibles and cheek teeth of U. rossicus from four localities of the South Siberia are examined. This species inhabited the steppe regions in early Middle and Late Pleistocene. By odontological characters it is more close to U. r. rossicus from Krasnodar, than to U. rossicus uralensis from Kizel Cave in Ural. Discriminant analysis, based on measurements of lower cheek teeth of the cave bears from seven sites of Europe and Siberia, demonstrated that U. rossicus most resembles morphometrically U. savini. As a result of cladistic analysis employed 17 characters of skull, limb bones, and dentition, the phylogenetic tree has been obtained for 7 species of the genus U r s u s. A four species of the cave bears are included in the subgenus Spelearctos: U. savini, U. rossicus, U. denin - geri and U. spelaeus. Key words: cave bears, U r s u s, Siberia, Pleistocene, evolution (1) Zoological Institute, Russian Academy of Sciences, Universitetskaya naberezhnaya 1, 199034 St. Petersburg, Russia; e-mail: bgf@zisp.spb.su (2) Institute of Geology, Siberian Branch of Russian Academy of Sciences, pr. Akad. Koptiuga 3, 630090 Novosibirsk, Russia; e-mail: foronova@uiggm.nsc.ru
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