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390 BARYSHNIKOV & FORONOVA CAD. LAB. XEOL. LAXE 26 (2001) Figure 12. Plot of centroids for each locality on the plane canonical variate 1 and 2 for m3. Table 9. Correlations of canonical variate for m3. demonstrates the largest m3 in three samples examined of U. rossicus (table 5), only the specimens from Krasnodar and Kizel Cave reliably differ in the greatest crown width (P
CAD. LAB. XEOL. LAXE 26 (2001) Pleistocene small cave Bear 391 Figure 13. Similarity dendrogram of samples of genus Ursus based on length and width means of lower cheek teeth p4, m1, m2 and m3. ples from Siberia and Krasnodar constitute one cluster, while the sample from Kizel Cave groups with that of U. savini. The obtained results of the discriminant analysis show the resemblance of U. rossicus with U. savini in proportions of the lower molars m1, m2 and m3. However in proportions of the lower premolar p4, the former species is more close to U. spelaeus. The enlargement and complication of p4 may be treated as a specialization of U. ros - sicus to consume rough herbaceous food. Postcranial skeleton The isolated limb bones of a small cave bear have been found in Mokhovo quarry and in Berezhekovo. To identify their sex we used the data on the size sex dimorphism in U. rossicus, established for the sample from Kizel Cave (see VERESHCHAGIN & BARYSHNIKOV in press). The specimens examined resemble in size the corresponding bones from Kizel Cave (table 10). The single fourth metacarpal, unlike those from Kizel Cave, does not have the incision on the distal articulate ridge. CLADISTIC ANALYSIS OF CAVE BEARS The subgenus Speleartos E.Geoffroy, 1833 includes four species: U. savini Andrews, 1922 (early Middle Pleistocene), U. deningeri von Reichenau, 1904 (Middle and Late Pleistocene), U. rossicus Borissiak, 1930 (Middle and Late Pleistocene) and U. spelaeus Rosenmüller, 1794 (late Middle and Late Pleistocene). The first and fourth species are found in Europe only, the both others are found in
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