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Report No. 6945<br />

BBN Systems <strong>and</strong> Technologies Corporation<br />

16 kHz (Table 2.7). Low to moderate frequency click sounds made by o<strong>the</strong>r<br />

species -have lower limits <strong>of</strong> 40 Hz to 2 kHz (Table 2.7).<br />

Echo-location clicks from too<strong>the</strong>d whales are <strong>the</strong> highest frequency sounds<br />

produced by any marine mammals. In <strong>the</strong> white whale <strong>the</strong>y range from 40 to 120<br />

kHz (Au e$ al. 1985, 1987) <strong>and</strong> in <strong>the</strong> harbor porpoise <strong>the</strong>y range from 100 to<br />

160 kHz (Mdhl <strong>and</strong> Anderson 1973). The sound intensity <strong>of</strong> echolocation clicks<br />

has been reported to range from 132 to 222 dB re 1 pPa at 1 m (Table 2.7).<br />

Moreover, <strong>the</strong>se signals are highly directional <strong>and</strong>, at least in <strong>the</strong> white<br />

whale <strong>and</strong> bottlenose dolphin, have beamwidths (to <strong>the</strong> -3, dB points) <strong>of</strong> 5 to 12<br />

degrees from <strong>the</strong> major axis (Au et al. 1986, 1987). Source levels <strong>and</strong><br />

frequencies are variable within as well as between species; too<strong>the</strong>d whales<br />

apparently adjust <strong>the</strong>ir click frequencies <strong>and</strong> levels for optimum echolocation<br />

capabilities under varying environmental conditions ( Au et al. 1985). When<br />

echolocating, <strong>the</strong> white whale usually emits a series <strong>of</strong> about 16 to 42 clicks<br />

(Au et al. 1985). In <strong>the</strong> white whale <strong>the</strong> typical interclick interval for <strong>the</strong><br />

main-echolocating clicks is 44 ms (Au et al. 1985). Typical click durations<br />

for odontocetes are less than' 1 ms <strong>and</strong> can be as low as 35 ps (Popper 1980).<br />

2.2.2 Baleen whales<br />

Most sounds made by fin, blue, minke <strong>and</strong> sei whales (genus ~alaeno~tera)<br />

are low in frequency <strong>and</strong> <strong>of</strong> moderate intensity. Most vocalizations are below<br />

3 kHz <strong>and</strong> have source levels <strong>of</strong> 15 1 to 188 dB re 1 pPa at 1 m (Table 2.8) .<br />

The fin whale produces a repeated stereotyped 20 Hz call during winter that<br />

could be a display associated with reproduction (Watkins 1981; Watkins et al.<br />

1987). .These calls have been recorded from most ice free waters in winter,<br />

but not specifically from <strong>the</strong> Bering Sea (Watkins et al. 1987). The<br />

significance <strong>and</strong> uses <strong>of</strong> o<strong>the</strong>r calls are unknown.<br />

Some moderate to high frequency click sounds have been recorded in <strong>the</strong><br />

presence <strong>of</strong> blue, fin <strong>and</strong> minke whales (Beamish <strong>and</strong> Mitchell 1971, 1973;<br />

Beamish 1979; Thompson et al. 1979). Frequencies were 3.3 to 31 kHz <strong>and</strong><br />

source levels were 130 to 159 dB re 1 pPa at 1 m (Table 2.8). Beamish <strong>and</strong><br />

Mitchell (1973) raised <strong>the</strong> possibility that baleen whales use echolocation.<br />

However, o<strong>the</strong>r researchers have not recorded <strong>the</strong>se click sounds <strong>and</strong> believe<br />

that <strong>the</strong>re is no evidence to show that baleen whales use echolocation (Norris<br />

1981 ; Watkins 1981).<br />

Humpbacks are very vocal when on <strong>the</strong>ir sou<strong>the</strong>rn wintering grounds. The<br />

songs <strong>and</strong> social sounds produced in late fall <strong>and</strong> winter have been well<br />

studied (Tyack 1981; Payne <strong>and</strong> Guinee 1983). Humpbacks do not sing <strong>and</strong> are<br />

less vocal when on <strong>the</strong>ir summering grounds in <strong>the</strong> Gulf <strong>of</strong> Alaska (Thompson et<br />

al. 1986). During summer, sounds are generally in <strong>the</strong> 20 to 2000 Hz range<br />

with intensities <strong>of</strong> 144 to 192 dB re 1 uPa at 1 m (Thompson et al. 1986).<br />

Gray whales are vocal when migrating <strong>and</strong> when on <strong>the</strong>ir sou<strong>the</strong>rn wintering<br />

grounds (Fish et al. 1974; Norris et al. 1977; Dahlheim et al. 1984). Sounds<br />

made on <strong>the</strong> summer feeding grounds in <strong>the</strong> Bering Sea are similar to those made<br />

while on <strong>the</strong> wintering grounds (Moore <strong>and</strong> Ljungblad 1984). The behavioral<br />

significance <strong>of</strong> <strong>the</strong> sounds is unknown (Dahlheim et al. 1984; Moore <strong>and</strong><br />

Ljungblad 1984). On both <strong>the</strong> summering <strong>and</strong> wintering grounds, frequencies <strong>of</strong>

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