Checklist of Bolivian Compositae - Royal Botanic Gardens, Kew

5. (3) Achenes glabrous; leaves 1–2.5 cm long; spines usually as long as or much longer than
leaves; c. 20 florets per capitulum
D. hystrix
Achenes setuliferous; leaves 4–8 (–10) cm long; spines short or absent; 11–20 florets per
capitulum
D. brasiliense [var. varians]
6. (2) Spines as long as or longer than leaves; phyllary apices with a short spine D. ferox
Spines short or absent; phyllary apices mucronate 7
7. (6) Corollas pubescent outside throughout D. inerme
Corollas glabrous except for lobe apices
D. brasiliense
8. (1) Adult leaves glabrous, lamina 50–100 20–60 mm; capitula 30–40 mm tall D. latifolium
Adult leaves with persistent pubescence beneath, lamina 25–70 10–35 mm; capitula 20–30
mm tall 9
9. (8) Florets 25–30; involucres 20–25 mm tall D. candolleanum
Florets 40–50; involucres 25–30 mm tall
D. velutinum
Dasyphyllum armatum (J. Koster) Cabrera, Revista Mus La Plata, Secc. Bot. 9: 51 (1959).
*Chuquiraga armata J. Koster, Blumea 5(3): 662 (1945). Type: [Bolivia:] ‗Hab.: Strauch an steinigen Abhängen
bei Pojos, 2400 m alt., April 1911, [Herzog] n. 2023.‘ Holotype: L(94437126); isotype: LP, S.
Flotovia armata (J. Koster) Tovar, Publ. Mus. Hist. Nat. ‗Javier Prado‘, Ser. B., Bot. 7: 7 (1953).
Argentina, Bolivia (Cochabamba).
Dry scrubby slopes.
2000–3000 m.
March–April.
Cochabamba: Wood 16012 (K).
Dasyphyllum brasiliense (Spreng.) Cabrera, Revista Mus. La Plata, Secc. Bot. 9: 72 (1959).
Joannea brasiliensis Spreng., Neue Entdeck. 2: 132 (1821). Type: ‗E Brasilia‘ [Sello]. (Lectotype, selected here, P;
isolectotype K). Sprengel‘s herbarium was divided and sold; the Compositae went to Schultz Bipontinus,
then to Herb. Cosson and thence to Paris (P) (Stafleu & Cowan 1985).
Carthamus fluminensis Vell., Fl. Flum. : 342 (1825)[7 Sept. - 28 Nov. 1829]; Fl. Flum. Icones 8: tab. 78 (1831).
Type: not cited.
Flotovia glabra Spreng., Syst. Veg., ed. 16, 3: 506 (1826), nom. illegit. superfl. pro Joannea brasiliensis Spreng.
Flotovia (Erinesia) quinquinervis Gardner, London J. Bot. 4: 127 (1845). Type: Brazil, Rio de Janeiro ‗Hab.
Woods, Organ Mountains, at an elevation of 3500 feet‘. July. [Gardner] 863. Lectotype (selected here): K – ex
Herb. Hookerianum; isotypes: BM, K – ex Herb. Benthamianum.
Chuquiraga glabra (Spreng.) Baker in Mart., Fl. Bras. 6(3): 363 (1884).
Chuquiraga racemosa Baker in Mart., Fl. Bras. 6(3): 363 (1884). Types: [Brazil:] ‗Habitat in prov. Minas Geraës
in sylvis apertis: Riedel n. 365; ad Canta Gallo: Peckolt n. 370‘. Lectotype (selected by Cabrera, 1959: 74):
Riedel n. 365, K; isolectoypes: G, LE, P.
*Chuquiraga brasiliensis (Spreng.) Kuntze, Revis. Gen. Pl. 3(3): 141 (1898).
Flotovia brasiliensis (Spreng.) Cabrera ex Tovar, Publ. Mus. Hist. Nat. ―Javier Prado‖, Ser. B., Bot. 7: 1 (1953).
Vernacular names: GUAIAPÁ-PARREIRA, CIPÓ-AGULHA (Cabrera & Klein, 1973).
var. barnadesioides (Tovar) Cabrera, Revista Mus. La Plata, Secc. Bot. 9: 76 (1959).
Flotovia barnadesioides Tovar, Publ. Mus. Hist. Nat. ―Javier Prado‖, Ser. B., Bot. 7: 10 (1953). Type: [Peru:]
‗Junin: Prov. Tarma, Colonia del Perené, 800-900 m s.m., leg. A. Weberbauer, 9-IX-1920‘. Holotype: USM;
isotype: LP (fragment).
Bolivia (Chuquisaca, La Paz, Santa Cruz), Peru.
700–1800 m.
September–January.
Chuquisaca: Wood 8457 (K).
var. divaricatum (Griseb.) Cabrera, Revista Mus. La Plata, Secc. Bot. 9: 77 (1959).
Barnadesia divaricata Griseb., Abh. Königl. Ges. Wiss. Göttingen 24(1): 209 (March-April 1879), Symb. Fl. Arg.
: 209 (1879). Types: ‗O.: Tarija, pr. Carapari, Cuesta de Buyuyu. (Paraguay: Bal[ansa] 822.)‘ Syntypes:
Balansa 822, Lorentz 627, Lorentz & Hieronymus 871, GOET. Lectotype (probably selected by Cabrera, 1959:
77, but as ‗Isotype‘): ‗Salta: Orán, Caraparì, leg. Hieronymus et Lorentz 627 (Isotypus: B; Fot. Field Mus.
195